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Genus Pseudomonas

Warning: In the List of Prokaryotic names with Standing in Nomenclature, an arrow () only indicates the sequence of valid publication of names and does not mean that the last name in the sequence must be used (see: Introduction).

Number of species cited in this file: 239
Number of subspecies cited in this file: 18

Classification (Warning: see also the file "Classification of prokaryotes: Introduction").


For a detailed description of this taxon see Bergey’s Manual of Systematics of Archaea and Bacteria (BMSAB).


 

Pseudomonas Migula 1894, genus. (Type genus of the order ¤ Pseudomonadales Orla-Jensen 1921 [Approved Lists 1980]; type genus of the suborder ¤ Pseudomonadineae Breed et al. 1957 [Approved Lists 1980]; type genus of the family ¤ Pseudomonadaceae Winslow et al. 1917 [Approved Lists 1980]; type genus of the tribe ¤ Pseudomonadeae Kluyver and Van Niel 1936 [Approved Lists 1980]).
Type species: ¤ Pseudomonas aeruginosa (Schroeter 1872) Migula 1900 (Approved Lists 1980).
Synonyms: "Chlorobacterium" Guillebeau 1890, nom. rejic. Opin. 6 (not "Chlorobacterium" Lauterborn 1916), "Liquidomonas" Orla-Jensen 1909, "Loefflerella" Holden 1935.
Etymology: Gr. adj.  pseudês, false; Gr. fem. n. monas, a unit, monad; N.L. fem. n. Pseudomonas, false monad.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MIGULA (W.): Über ein neues System der Bakterien. Arbeiten aus dem Bakteriologischen Institut der Technischen Hochschule zu Karlsruhe, 1894, 1, 235-238.

Notes:
1 Pseudomonas Migula 1894 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Chryseomonas Holmes et al. 1986, and an earlier heterotypic synonym of ¤ Flavimonas Holmes et al. 1987.
Reference: ANZAI (Y.), KUDO (Y.) and OYAIZU (H.): The phylogeny of the genera Chryseomonas, Flavimonas, and Pseudomonas supports synonymy of these three genera. Int. J. Syst. Bacteriol., 1997, 47, 249-251.
Original article in IJSEM Online
2 According to Youg and Park 2007, Pseudomonas Migula 1894 (Approved Lists 1980) is a probable earlier heterotypic synonym of ¤ Azotobacter Beijerinck 1901 (Approved Lists 1980) [which is a probable earlier heterotypic synonym of ¤ Azomonas Winogradsky 1938 (Approved Lists 1980)].
Reference: YOUNG (J.M.) and PARK (D.C.): Probable synonymy of the nitrogen-fixing genus Azotobacter and the genus Pseudomonas. Int. J. Syst. Evol. Microbiol., 2007, 57, 2894-2901.
Original article in IJSEM Online

Pseudomonas Migula 1894 emend. Yang et al. 2013.
Reference: YANG, G., HAN, L., WEN, J. and ZHOU, S. 2013. Pseudomonas guangdongensis sp. nov., isolated from an electroactive biofilm, and emended description of the genus Pseudomonas Migula 1894. Int. J. Syst. Evol. Microbiol., 63, 4599-4605.
Original article in IJSEM Online

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Pseudomonas abietaniphila Mohn et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) BKME-9 = ATCC 700689= CCUG 50779 = CIP 106708.
Sequence accession no. (16S rRNA gene) for the type strain: AJ011504.
Etymology: N.L. neut. n. abietanum, abietane; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. abietaniphila, abietane-loving.
Valid publication: Validation List no. 70. Int. J. Syst. Bacteriol., 1999, 49, 935-936.
Effective publication: MOHN (W.W.), WILSON (A.E.), BICHO (P.) and MOORE (E.R.B.): Physiological and phylogenetic diversity of bacteria growing on resin acids. Syst. Appl. Microbiol., 1999, 22, 68-78.
Validation List no. 70 in IJSEM Online - Effective publication Online

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Pseudomonas acidovorans den Dooren de Jong 1926, species.
Type strain: (see also StrainInfo.net) KS 0057 = Stanier 14 = den Dooren de Jong 7 = ACM 489 = ATCC 15668= CCUG 12692 = CCUG 14481 = CFBP 2444 = CIP 103021 = DSM 39 = IAM 12409 = IFO (now NBRC) 14950 = JCM 5833 = JCM 6203 = LMG 1226 = LMG 6031 = NCCB 26037 = NCIB (now NCIMB) 9681 = NCTC 10683.
Sequence accession no. (16S rRNA gene) for the type strain: AB021417.
Synonyms: "Pseudomonas indoloxidans" Gray 1928, "Pseudomonas desmolytica" Gray and Thornton 1928.
Etymology: N.L. neut. n. acidum (from L. adj. acidus -a -um, sour), an acid; L. v. voro, to eat, to devour; N..L. part. adj. acidovorans, acid devouring.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DEN DOOREN DE JONG (L.E.): Bijdrage tot de kennis van het Mineralisatieproces. Thesis. Technische Hogeschool, Delft, 1926, pp. 1-199.

¤ Comamonas acidovorans (den Dooren de Jong 1926) Tamaoka et al. 1987, comb. nov.

¤ Delftia acidovorans (den Dooren de Jong 1926) Wen et al. 1999, comb. nov.

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Pseudomonas aeruginosa (Schroeter 1872) Migula 1900, species. (Type species of the genus.)
Type strain: (see also StrainInfo.net) RH 815 = ATCC 10145= ATCC 10145-U = CCEB 481 = CCUG 551 = CCUG 28447 = CCUG 29297 = CFBP 2466 = CIP 100720 = DSM 50071 = IBCS 277 = IFO (now NBRC) 12689 = JCM 5962 = LMG 1242 = NCCB 76039 = NCIB (now NCIMB) 8295 = NCTC 10332 = NRRL B-771 = VKM B-588.
Sequence accession no. (16S rRNA gene) for the type strain: HE978271.
Synonyms: "Bacterium aeruginosum" Schroeter 1872, "Bacterium aeruginosum" Cohn 1872, "Micrococcus pyocyaneus" Zopf 1884, "Bacillus aeruginosus" (Schroeter 1872) Trevisan 1885, "Bacillus pyocyaneus" (Zopf 1884) Flügge 1886, "Pseudomonas pyocyanea" (Zopf 1884) Migula 1895, "Bacterium pyocyaneum" (Zopf 1884) Lehmann and Neumann 1896, probable synonym or variety: "Pseudomonas polycolor" Clara 1930.
Etymology: L. fem. adj. aeruginosa, full of copper rust, verdigris, hence green.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SCHROETER (J.): Über einige durch Bacterien gebildete Pigmente. (1872). In: F. COHN (ed.), Beitrage zur Biologie der Pflanzen, J.U. Kern's Verlag, Breslau, 1875, pp. 109-126. 2 MIGULA (W.): System der Bakterien, Vol. 2, Gustav Fischer, Jena, 1900.

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Pseudomonas aestusnigri Sanchez et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) VGXO14=CCUG 64165=CECT 8317.
Sequence accession no. (16S rRNA) for the type strain: HG004394.
Whole-genome sequence accession no. for the type strain: -.
Etymology: -.
Source: Environmental.
Valid publication: Validation List no. 159. List of new names and new combinations previously effectively, but not validly, published. Int. J Syst. Evol. Microbiol., 64 (2014), 2927-2929.
Validation List Online
Effective publication: SÁNCHEZ, D., MULET, M., RODRÍGUEZ, A. C., DAVID, Z., LALUCAT, J. and GARCÍA-VALDÉS, E. 2014. Pseudomonas aestusnigri sp. nov., isolated from crude oil-contaminated intertidal sand samples after the Prestige oil spill. Syst. Appl. Microbiol., 37, 89-94
Effective publication Online

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Pseudomonas agarici Young 1970, species.
Type strain: (see also StrainInfo.net) ATCC 25941= CCUG 32769 = CFBP 2063 = CIP 106703 = DSM 11810 = ICMP 2656 = JCM 12566 = LMG 2112 = NCPPB 2289.
Sequence accession no. (16S rRNA gene) for the type strain: Z76652.
Etymology: N.L. n. Agaricus, a genus of fungi; N.L. gen. n. agarici, of Agaricus.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: YOUNG (J.M.): Drippy gill: a bacterial disease of cultivated mushrooms caused by Pseudomonas agarici n. sp. New Zealand Journal of Agricultural Research, 1970, 13, 977-990.

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Pseudomonas alcaligenes Monias 1928, species.
Type strain: (see also StrainInfo.net) ATCC 14909= CCUG 1425 A = CFBP 2437 = CIP 101034 = DSM 50342 = IFO (now NBRC) 14159 = JCM 5967 = LMG 1224 = NCCB 76044 = NCTC 10367 = VKM B-2171.
Sequence accession no. (16S rRNA gene) for the type strain: D84006.
Etymology: N.L. n. alcali (from Arabic article al, the; Arabic n. qaliy, ashes of saltwort), alkali; N.L. suff. -genes (from Gr. v. gennaô, to produce), producing; N.L. adj. alcaligenes, alkali-producing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MONIAS (B.L.): Classification of Bacterium alcaligenes pyocyaneum and fluorescens. Journal of Infectious Diseases, 1928, 43, 330-334.

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Pseudomonas alcaliphila Yumoto et al. 2001, sp. nov.
Type strain: (see also StrainInfo.net) AL15-21 = IAM 14884 = JCM 10630 = NBRC 102411.
Sequence accession no. (16S rRNA gene) for the type strain: AB030583.
Etymology: N.L. n. alcali (from Arabic article al, the; Arabic n. qaliy, ashes of saltwort), alcali; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. alcaliphila, liking alkaline environments.
Valid publication: YUMOTO (I.), YAMAZAKI (K.), HISHINUMA (M.), NODASAKA (Y.), SUEMORI (A.), NAKAJIMA (K.), INOUE (N.) and KAWASAKI (K.): Pseudomonas alcaliphila sp. nov., a novel facultatively psychrophilic alkaliphile isolated from seawater. Int. J. Syst. Evol. Microbiol. 2001, 51, 349-355.
Original article in IJSEM Online

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Pseudomonas alkylphenolica Mulet et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) KL28=JCM 16553=KCTC 22206.
Sequence accession no. (complete genome) for the type strain: CP009048.
Whole-genome sequence accession no. for the type strain: CP009048.
Etymology: N.L. fem. adj. alkylphenolica, pertaining to alkylphenol compounds.
Source: Environmental - soil.
Valid publication: MULET, M., SÁNCHEZ, D., LALUCAT, J., LEE, K. and GARCÍA-VALDÉS, E. 2015. Pseudomonas alkylphenolica sp. nov., a bacterial species able to form special aerial structures when grown on p-cresol. Int. J. Syst. Evol. Microbiol., 65, 4013-4018.
Original article in IJSEM Online

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Pseudomonas aminovorans den Dooren de Jong 1926, species.
Type strain: (see also StrainInfo.net) ATCC 23314= CCUG 2081 = CIP 106737 = DSM 7048 = JCM 7852 = LMG 2122 = NCCB 26039 = NCIB (now NCIMB) 9039 = NCTC 10684 = VKM B-2058.
Sequence accession no. (16S rRNA gene) for the type strain: AJ011759.
Etymology: N.L. n. aminum, amine; L. v. voro, to eat, to devour; N.L. part. adj. aminovorans, amine-devouring, digesting.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DEN DOOREN DE JONG (L.E.): Bijdrage tot de kennis van het Mineralisatieproces. Thesis. Technische Hogeschool, Delft. pp. 1-199.

¤ Aminobacter aminovorans (den Dooren de Jong 1926) Urakami et al. 1992, comb. nov.

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Pseudomonas amygdali Psallidas and Panagopoulos 1975, species.
Type strain: (see also StrainInfo.net) ATCC 33614= CCUG 32770 = CFBP 3205 = CIP 106734 = DSM 7298 = ICMP 3918 = LMG 2123 = LMG 13184 = NCPPB 2607.
Sequence accession no. (16S rRNA gene) for the type strain: Z76654.
Etymology: L. gen. n. amygdali, of the almond.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: PSALLIDAS (P.G.) and PANAGOPOULOS (C.G.): A new bacteriosis of almond caused by Pseudomonas amygdali sp. nov. Annales de l'Institut Phytopathologique Benaki N.S., 1975, 11, 94-108.

Note: Pseudomonas amygdali Psallidas and Panagopoulos 1975 (Approved Lists 1980), ¤ Pseudomonas ficuserectae Goto 1983, ¤ Pseudomonas meliae Ogimi 1981, and ¤ Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992 are members of a single genomospecies. Thus, these species should be considered synonymous and the correct name for this species should be Pseudomonas amygdali. Unfortunately, Pseudomonas amygdali, as delineated by DNA-DNA hybridization, could not be differentiated from other genomospecies by phenotypic tests. So the authors do not formally propose nomenclatural changes.
Reference: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Šutič and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

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Pseudomonas andropogonis (Smith 1911) Stapp 1928, species.
Type strain: (see also StrainInfo.net) ATCC 23061= CCUG 32772 = CFBP 2421 = CIP 105771 = DSM 9511 = ICMP 2807 = JCM 10487 = LMG 2129 = NCPPB 934 = NRRL B-14296.
Sequence accession no. (16S rRNA gene) for the type strain: X67037.
Synonyms: "Bacterium andropogoni" (sic) Smith 1911, "Aplanobacter stizolobii" Wolf 1920, "Pseudomonas stizolobii" (Wolf 1920) Stapp 1935.
Etymology: N.L. n. Andropogon (from Gr. n. aner andros, man; Gr. pogon pogonis, beard), a botanical generic name of widely distributed grasses; N.L. gen. n. andropogonis, of Andropogon.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SMITH (E.F.): Bacteria in relation to plant diseases. Carnegie Institute, Washington, 1911, 2, 1-368. 2 STAPP (C.): Schizomycetes (Spaltpilze oder Bakterien). In: SORAUER (ed.), Handbuch der Pflanzenkrankheiten, 5th ed., Vol. 2, Paul Parey, Berlin, 1928, pp. 1-295.

Note: On the Approved Lists of Bacterial Names and on the Approved Lists of Bacterial Names (Amended Edition), the page numbers of the paper by Stapp are cited as pp. 758.

¤ Burkholderia andropogonis (Smith 1911) Gillis et al. 1995, comb. nov.

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Pseudomonas anguilliseptica Wakabayashi and Egusa 1972, species.
Type strain: (see also StrainInfo.net) ATCC 33660= CCUG 35503 = CIP 106711 = DSM 12111 = NCIMB 1949.
Sequence accession no. (16S rRNA gene) for the type strain: X99540.
Etymology: L. n. anguilla, eel; L. fem. adj. septica, producing a putrefaction, putrefying, septic; N.L. fem. adj. anguilliseptica, pertaining to diseased eels.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: WAKABAYASHI (H.) and EGUSA (S.): Characteristics of a Pseudomonas sp. from an epizootic of pond-cultured eels (Anguilla japonica). Bulletin of the Japanese Society of Scientific Fisheries, 1972, 38, 577-587.

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Pseudomonas antarctica Reddy et al. 2004, sp. nov.
Type strain: (see also StrainInfo.net) CMS 35 = CCUG 49625 = DSM 15318 = MTCC 4992.
Sequence accession no. (16S rRNA gene) for the type strain: AJ537601.
Etymology: L. fem. adj. antarctica, southern, Antarctic.
Valid publication: REDDY (G.S.N.), MATSUMOTO (G.I.), SCHUMANN (P.), STACKEBRANDT (E.) and SHIVAJI (S.): Psychrophilic pseudomonads from Antarctica: Pseudomonas antarctica sp. nov., Pseudomonas meridiana sp. nov. and Pseudomonas proteolytica sp. nov. Int. J. Syst. Evol. Microbiol., 2004, 54, 713-719.
Original article in IJSEM Online

Note: The specific epithet is a L. fem. adj., not a N.L. fem. adj. as cited in the paper by Reddy et al. 2004.

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Pseudomonas antimicrobica Attafuah and Bradbury 1990, sp. nov.
Type strain: (see also StrainInfo.net) NCIB (now NCIMB) 9898 = LMG 18920.
Sequence accession no. (16S rRNA gene) for the type strain: AB021384.
Etymology: Gr. pref. anti, over against; N.L. neut. n. microbium (from Gr. adj. mikros, small and Gr. n. bios, life), a microbe; L. fem. suff. -ica, suffix used with the sense of pertaining to; N.L. fem. adj. antimicrobica, against microbes (referring to a wide antimicrobial activity).
Valid publication: Validation List no. 34. Int. J. Syst. Bacteriol., 1990, 40, 320-321.
Effective publication: ATTAFUAH (A.) and BRADBURY (J.F.): Pseudomonas antimicrobica, a new species strongly antagonistic to plant pathogens. J. Appl. Bacteriol., 1989, 67, 567-573.
Validation List no. 34 in IJSEM Online

¤ Burkholderia gladioli (Severini 1913) Yabuuchi et al. 1993, comb. nov.

Note: According to Coenye et al. 2000, Pseudomonas antimicrobica Attafuah and Bradbury 1990 is a later heterotypic synonym of Burkholderia gladioli (Severini 1913) Yabuuchi et al. 1993, comb. nov.
Reference: COENYE (T.), GILLIS (M.) and VANDAMME (P.): Pseudomonas antimicrobica Attafuah and Bradbury 1990 is a junior synonym of Burkholderia gladioli (Severini 1913) Yabuuchi et al. 1993. Int. J. Syst. Evol. Microbiol. 2000, 50, 2135-2139.
Original article in IJSEM Online

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Pseudomonas argentinensis Peix et al. 2005, sp. nov.
Type strain: (see also StrainInfo.net) CH01 = CECT 7010 = LMG 22563.
Sequence accession no. (16S rRNA gene) for the type strain: AY691188.
Etymology: N.L. fem. adj. argentinensis, pertaining to the Argentine, of the Argentine.
Valid publication: PEIX (A.), BERGE (O.), RIVAS (R.), ABRIL (A.) and VELÁZQUEZ (E.): Pseudomonas argentinensis sp. nov., a novel yellow pigment-producing bacterial species, isolated from rhizospheric soil in Córdoba, Argentina. Int. J. Syst. Evol. Microbiol., 2005, 55, 1107-1112.
Original article in IJSEM Online

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Pseudomonas arsenicoxydans Campos et al. 2011, sp. nov.
Type strain: (see also StrainInfo.net) VC-1 = CCUG 58201 = CECT 7543.
Sequence accession no. (16S rRNA gene) for the type strain: FN645213.
Etymology: L. n. arsenicum, arsenic; N.L. part. adj. oxydans, oxidizing; N.L. part. adj. arsenicoxydans, arsenic-oxidizing.
Valid publication: Validation List no. 137. Int. J. Syst. Evol. Microbiol., 2011, 61, 1-3.
Effective publication: CAMPOS (V.L.), VALENZUELA (C.), YARZA (P.), KÄMPFER (P.), VIDAL (R.), ZAROR (C.), MONDACA (M.A.), LOPEZ-LOPEZ (A.) and ROSSELLÓ-MÓRA (R.): Pseudomonas arsenicoxydans sp nov., an arsenite-oxidizing strain isolated from the Atacama desert. Syst. Appl. Microbiol., 2010, 33, 193-197.
Validation List no. 137 in IJSEM Online - Effective publication Online

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Pseudomonas asplenii (Ark and Tompkins 1946) Savulescu 1947, species.
Type strain: (see also StrainInfo.net) ATCC 23835= CCUG 32773 = CFBP 3279 = CIP 106710 = DSM 17133 = ICMP 3944 = LMG 2137 = NCPPB 1947.
Sequence accession no. (16S rRNA gene) for the type strain: AB021397.
Synonym: "Phytomonas asplenii" Ark and Tompkins 1946.
Etymology: N.L. n. Asplenium, genus of ferns, spleenworts; N.L. gen. n. asplenii, of Asplenium.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 ARK (P.A.) and TOMPKINS (C.M.): Bacterial leaf blight of bird's-nest fern. Phytopathology, 1946, 36, 758-761. 2 SAVULESCU (T.): Contribution à la classification des bacteriacées phytopathogènes. Analele Academiei Romane Series III Tom 22, Memoire, 1947, 4, 1-26.

Pseudomonas asplenii (Ark and Tompkins 1946) Savulescu 1947 (Approved Lists 1980) emend. Tvrzová et al. 2006.
Type strain: (see also StrainInfo.net) ATCC 23835= CCUG 32773 = CFBP 3279 = CIP 106710 = DSM 17133 = ICMP 3944 = LMG 2137 = NCPPB 1947.
Sequence accession no. (16S rRNA gene) for the type strain: AB021397.
Synonym: "Phytomonas asplenii" Ark and Tompkins 1946.
Etymology: N.L. n. Asplenium, genus of ferns, spleenworts; N.L. gen. n. asplenii, of Asplenium.
References: TVRZOVÁ (L.), SCHUMANN (P.), SPRÖER (C.), SEDLÁČEK (I.), PÁČOVÁ (Z.), ŠEDO (O.), ZDRÁHAL (Z.), STEFFEN (M.) and LANG (E.): Pseudomonas moraviensis sp. nov. and Pseudomonas vranovensis sp. nov., soil bacteria isolated on nitroaromatic compounds, and emended description of Pseudomonas asplenii. Int. J. Syst. Evol. Microbiol., 2006, 56, 2657-2663.
Original article in IJSEM Online

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Pseudomonas asturiensis González et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) LPPA 221=LMG 26898=CECT 8095.
Sequence accession no. (16S rRNA) for the type strain: FM865870.
Etymology: N.L. fem. adj. asturiensis, from Asturias, where the strains were isolated.
Source: Plant.
Valid publication: Validation List no. 154. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 63 (2013), 3931-3934.
Validation List Online
Effective publication: GONZÁLEZ, A. J., CLEENWERCK, I., DE VOS, P. and FERNÁNDEZ-SANZ, A. M. 2013. Pseudomonas asturiensis sp. nov., isolated from soybean and weeds. Syst. Appl. Microbiol., 36, 320-324.
Effective publication Online

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Pseudomonas asuensis Reddy and Garcia-Pichel 2015, sp. nov.
Type strain: (see also StrainInfo.net) CP 155-2=LMG 28687=KCTC 32484.
Sequence accession no. (16S rRNA) for the type strain: AJ871471.
Whole-genome sequence accession no. for the type strain: -.
Etymology: N.L. fem. adj. asuensis, an adjective arbitrarily derived from Arizona State University.
Source: Environmental - soil.
Valid publication: Validation List no. 164. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 65 (2015), 2017-2025.
Validation List Online
Effective publication: Reddy, G.S. and Garcia-Pichel, F. 2015. Description of Pseudomonas asuensis sp. nov. from biological soil crusts in the Colorado plateau, United States of America. J. Microbiol., 53, 6-13.
Effective publication Online

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Pseudomonas aurantiaca Nakhimovskaya 1948, species.
Type strain: (see also StrainInfo.net) ATCC 33663= CIP 106718 = NCIB (now NCIMB) 10068 = VKM B-876.
Sequence accession no. (16S rRNA gene) for the type strain: DQ682655.
Etymology: N.L. fem. adj. aurantiaca, orange colored.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: NAKHIMOVSKAYA (M.I.): Pseudomonas aurantiaca n. sp. Mikrobiologiya, 1948, 17, 58-65.

¤ Pseudomonas chlororaphis subsp. aurantiaca (Nakhimovskaya 1948) Peix et al. 2007, comb. nov.

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Pseudomonas aureofaciens Kluyver 1956, species.
Type strain: (see also StrainInfo.net) ATCC 13985= CFBP 2133 = DSM 6698 = LMG 1245 = NCIMB 9030.
Sequence accession no. (16S rRNA gene) for the type strain: AY509898.
Etymology: L. adj. aureus, golden; L. part. adj. faciens, making; N.L. part. adj. aureofaciens, making golden.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: KLUYVER (A.J.): Pseudomonas aureofaciens nov. spec. and its pigments. Journal of Bacteriology, 1956, 72, 406-411.

¤ Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930 (Approved Lists 1980).

Note: According to Johnson and Palleroni 1989, Pseudomonas aureofaciens Kluyver 1956 (Approved Lists 1980) is a later heterotypic synonym of Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930 (Approved Lists 1980).
Reference: JOHNSON (J.L.) and PALLERONI (N.J.): Deoxyribonucleic acid similarities among Pseudomonas species. Int. J. Syst. Bacteriol., 1989, 39, 230-235.
Original article in IJSEM Online

¤ Pseudomonas chlororaphis subsp. aureofaciens (Kluyver 1956) Peix et al. 2007, comb. nov.

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Pseudomonas avellanae Janse et al. 1997, sp. nov.
Type strain: (see also StrainInfo.net) F11 = CIP 105176 = DSM 11809 = JCM 11937 = NCPPB 3487.
Sequence accession no. (16S rRNA gene) for the type strain: not found.
Etymology: N.L. gen. avellanae, of avellana, the species name of the host, Corylus avellana (hazlnut).
Valid publication: Validation List no. 61. Int. J. Syst. Bacteriol., 1997, 47, 601-602.
Effective publication: JANSE (J.D.), ROSSI (P.), ANGELUCCI (L.), SCORTICHINI (M.), DERKS (J.H.J.), AKKERMANS (A.D.L.), DE VRIJER (R.) and PSALLIDAS (P.G.): Reclassification of Pseudomonas syringae pv. avellanae as Pseudomonas avellanae (spec. nov.), the bacterium causing canker of hazelnut (Corylus avellana L.) Syst. Appl. Microbiol., 1996, 19, 589-595.
Validation List no. 61 in IJSEM Online - Effective publication Online

Note: Pseudomonas avellanae was previously known as Pseudomonas syringae pv. Avellanae.

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Pseudomonas avenae Manns 1909, species.
Type strain: (see also StrainInfo.net) ATCC 19860= CCUG 15838 = CFBP 2425 = CIP 106500 = DSM 7227 = ICMP 3183 = JCM 20985 = LMG 2117 = NCPPB 1011.
Sequence accession no. (16S rRNA gene) for the type strain: CP002521 (complete genome).
Etymology: N.L. n. Avena, genus of plants; N.L. gen. n. avenae, of Avena.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MANNS (T.F.): The blade blight of oats, a bacterial disease. Bulletin of the Ohio Agriculture Experiment Station, 1909, 210, 91-167.

Note: According to Hu et al. 1991, Pseudomonas avenae Manns 1909 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas rubrilineans (Lee et al. 1925) Stapp 1928 (Approved Lists 1980) and an earlier heterotypic synonym of ¤ Pseudomonas cattleyae (Pavarino 1911) Savulescu 1947 (Approved Lists 1980).
Reference: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

¤ Acidovorax avenae (Manns 1909) Willems et al. 1992, comb. nov.

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Pseudomonas avenae subsp. avenae (Manns 1909) Hu et al. 1991, subsp. nov.
Type strain: (see also StrainInfo.net) ATCC 19860= CCUG 15838 = CFBP 2425 = CIP 106500 = DSM 7227 = ICMP 3183 = JCM 20985 = LMG 2117 = NCPPB 1011.
Sequence accession no. (16S rRNA gene) for the type strain: AF137505.
Etymology: N.L. n. Avena, genus of plants; N.L. gen. n. avenae, of Avena.
Valid publication: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

Notes:
1 The original authorship, Manns 1909, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online
2 The subspecies name Pseudomonas avenae subsp. avenae (Manns 1909) Hu et al. 1991 is automatically created by the valid publication of Pseudomonas avenae subsp. citrulli (Schaad et al. 1978) Hu et al. 1991 and the valid publication of Pseudomonas avenae subsp. konjaci (Goto 1983) Hu et al. 1991 [Rule 40d (formerly Rule 46)].

¤ Acidovorax avenae subsp. avenae (Manns 1909) Willems et al. 1992, comb. nov.

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Pseudomonas avenae subsp. citrulli (Schaad et al. 1978) Hu et al. 1991, comb. nov.
Type strain: (see also StrainInfo.net) FC-247 = ATCC 29625= CCUG 17393 = CFBP 4459 = CIP 106436 = IBSBF 1851 = ICMP 7500 = ICPB 30064 = LMG 5376 = NCPPB 3679.
Sequence accession no. (16S rRNA gene) for the type strain: AF137506.
Basonym: ¤ Pseudomonas pseudoalcaligenes subsp. citrulli Schaad et al. 1978 (Approved Lists 1980).
Etymology: N.L. n. Citrullus, a genus of melon plants; N.L. gen. n. citrulli, of Citrullus.
Valid publication: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

¤ Acidovorax avenae subsp. citrulli (Schaad et al. 1978) Willems et al. 1992, comb. nov.

¤ Acidovorax citrulli (Schaad et al. 1978) Schaad et al. 2009, comb. nov.

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Pseudomonas avenae subsp. konjaci (Goto 1983) Hu et al. 1991, comb. nov.
Type strain: (see also StrainInfo.net) K2 = ATCC 33996= CCUG 17394 = CFBP 4460 = CIP 106439 = DSM 7481 = ICMP 7733 = JCM 2397 = LMG 5691 = NCPPB 3698 = PDDCC 7733.
Sequence accession no. (16S rRNA gene) for the type strain: AJ420325.
Basonym: ¤ Pseudomonas pseudoalcaligenes subsp. konjaci Goto 1983.
Etymology: N.L. gen. n. konjaci, of the konjac plant, Amorphophallus rivieri cv. Konjac.
Valid publication: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

¤ Acidovorax konjaci (Goto 1983) Willems et al. 1992, comb. nov.

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Pseudomonas azotifigens Hatayama et al. 2005, sp. nov.
Type strain: (see also StrainInfo.net) 6H33b = ATCC BAA-1049 = JCM 12708.
Sequence accession no. (16S rRNA gene) for the type strain: AB189452.
Etymology: N.L. n. azotum (from Fr. n. azote), nitrogen; L. part. adj. figens, fixing; N.L. part. adj. azotifigens, nitrogen fixing.
Valid publication: HATAYAMA (K.), KAWAI (S.), SHOUN (H.), UEDA (Y.) and NAKAMURA (A.): Pseudomonas azotifigens sp. nov., a novel nitrogen-fixing bacterium isolated from a compost pile. Int. J. Syst. Evol. Microbiol., 2005, 55, 1539-1544.
Original article in IJSEM Online

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Pseudomonas azotoformans Iizuka and Komagata 1963, species.
Type strain: (see also StrainInfo.net) CCUG 12536 = CIP 106744 = IAM 1603 = IFO (now NBRC) 12693 = JCM 2777.
Sequence accession no. (16S rRNA gene) for the type strain: D84009.
Etymology: N.L. n. azotum (from Fr. n. azote), nitrogen; L. v. formo, to fashion, to form; N.L. part. adj. azotoformans, nitrogen forming (by denitrification).
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IIZUKA (H.) and KOMAGATA (K.): New species of Pseudomonas belonged to fluorescent group (Studies on the microorganisms of cereal grains. Part V). Journal of the Agricultural Chemical Society of Japan, 1963, 37, 137-141.

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Pseudomonas baetica López et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) a390 = CECT 7720 = LMG 25716.
Sequence accession no. (16S rRNA gene) for the type strain: FM201274.
Etymology: L. fem. adj. baetica, of or belonging to Hispania Baetica, the name of the former province of the Roman Empire covering the south-west region of Spain.
Valid publication: LÓPEZ (J.R.), DIÉGUEZ (A.L.), DOCE (A.), DE LA ROCA (E.), DE LA HERRAN (R.), NAVAS (J.I.), TORANZO (A.E.) and ROMALDE (J.L.): Pseudomonas baetica sp. nov., a fish pathogen isolated from wedge sole, Dicologlossa cuneata (Moreau). Int. J. Syst. Evol. Microbiol., 2012, 62, 874-882.
Original article in IJSEM Online

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Pseudomonas balearica Bennasar et al. 1996, sp. nov.
Type strain: (see also StrainInfo.net) SP1402 = CCUG 44487 = CIP 105297 = DSM 6083.
Sequence accession no. (16S rRNA gene) for the type strain: U26418.
Etymology: L. fem. adj. balearica, of the Balearic Islands, where the organism was isolated.
Valid publication: BENNASAR (A.), ROSSELLÓ-MORA (R.), LALUCAT (J.) and MOORE (E.R.B.): 16 rRNA gene sequence analysis relative to genomovars of Pseudomonas stutzeri and proposal of Pseudomonas balearica sp. nov. Int. J. Syst. Bacteriol., 1996, 46, 200-205.
Original article in IJSEM Online

Note: Pseudomonas balearica accomodates the ¤ Pseudomonas stutzeri genomovar 6 strains.

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Pseudomonas bauzanensis Zhang et al. 2011, sp. nov.
Type strain: (see also StrainInfo.net) BZ93 = CGMCC 1.9095 = DSM 22558 = LMG 26048.
Sequence accession no. (16S rRNA gene) for the type strain: GQ161991.
Etymology: N.L. fem. adj. bauzanensis, of or belonging to Bauzanum medieval Latin name of Bozen/Bolzano, a city in South Tyrol, Italy, where the species was first isolated.
Valid publication: ZHANG (D.C.), LIU (H.C.), ZHOU (Y.G.), SCHINNER (F.) and MARGESIN (R.): Pseudomonas bauzanensis sp. nov., isolated from soil. Int. J. Syst. Evol. Microbiol., 2011, 61, 2333-2337.
Original article in IJSEM Online

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Pseudomonas beijerinckii Hof 1935, species.
Type strain: (see also StrainInfo.net) ATCC 19372= CCUG 49679 = CIP 106957 = DSM 7218 = JCM 13305 = JCM 21422 = LMG 2148 = NCCB 35008 = NCIMB 9041 = NRRL B-3153.
Sequence accession no. (16S rRNA gene) for the type strain: AB021386.
Etymology: N.L. gen. masc. n. beijerinckii, of Beijerinck, named after M.W. Beijerinck.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HOF (T.): An investigation of the microorganisms commonly present in salted beans. Recueil des Travaux Botaniques Néerlandais, 1935, 32, 151-173.

Note: According to Anzai et al., Pseudomonas beijerinckii Hof 1935 (Approved Lists 1980) does not belong to the genus Pseudomonas sensu stricto [1]. In 2006, Peçonek et al. propose the new combination ¤ Chromohalobacter beijerinckii (Hof 1935) Peçonek et al. 2006 [2] (see below).
References:
1 ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.
Original article in IJSEM Online
2 PEÇONEK (J.), GRUBER (C.), GALLEGO (V.), VENTOSA (A.), BUSSE (H.J.), KÄMPFER (P.), RADAX (C.) and STAN-LOTTER (H.): Reclassification of Pseudomonas beijerinckii Hof 1935 as Chromohalobacter beijerinckii comb. nov., and emended description of the species. Int. J. Syst. Evol. Microbiol., 2006, 56, 1953-1957.
Original article in IJSEM Online

¤ Chromohalobacter beijerinckii (Hof 1935) Peçonek et al. 2006, comb. nov.

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Pseudomonas benzenivorans Lang et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) CIP 109857 = DSM 8628.
Sequence accession no. (16S rRNA gene) for the type strain: FM208263.
Etymology: N.L. n. benzenum, benzene; L. part. adj. vorans, devouring, eating; N.L. part. adj. benzenivorans, benzene-devouring.
Valid publication: Validation List no. 145. Int. J. Syst. Evol. Microbiol., 2012, 62, 1017-1019.
Effective publication: LANG (E.), BURGHARTZ (M.), SPRING (S.), SWIDERSKI (J.) and SPRÖER (C.): Pseudomonas benzenivorans sp. nov. and Pseudomonas saponiphila sp. nov., represented by xenobiotics degrading type strains. Curr. Microbiol., 2010, 60, 85-91.
Validation List no. 145 in IJSEM Online - Effective publication Online

Pseudomonas benzenivorans Lang et al. 2012 emend. Hameed et al. 2014.
Reference: HAMEED, A., SHAHINA, M., LIN, S.-Y., LIU, Y.-C. and YOUNG, C.-C. 2014. Pseudomonas hussainii sp. nov., isolated from droppings of a seashore bird, and emended descriptions of Pseudomonas pohangensis, Pseudomonas benzenivorans and Pseudomonas segetis. Int. J. Syst. Evol. Microbiol., 64, 2330-2337.
Original article.

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Pseudomonas beteli corrig. (Ragunathan 1928) Savulescu 1947, species.
Type strain: (see also StrainInfo.net) ATCC 19861= CFBP 4337 = ICMP 2820 = LMG 978 = NCPPB 323.
Sequence accession no. (16S rRNA gene) for the type strain: AB021406.
Synonym: "Bacterium betle" Ragunathan 1928.
Etymology: N.L. n. betelum (from Malayan noun betle), betel; N.L. gen. n. beteli, of betel.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 RAGUNATHAN (C.): Bacterial leaf spot of betel. Annals of the Royal Botanical Garden of Peradeniya, Ceylon, 1928, 11, 51-61. 2 SAVULESCU (T.): Contribution à la classification des bacteriacées phytopathogènes. Analele Academiei Romane Series III, Tom 22, Memoire, 1947, 4, 1-26.

Notes:
1 The original spelling of the specific epithet, betle (sic), has been corrected by Trüper and De'Clari 1997.
Reference: TRÜPER (H.G.) and DE' CLARI (L.): Taxonomic note: Necessary correction of specific epithets formed as substantives (nouns) "in apposition". Int. J. Syst. Bacteriol., 1997, 47, 908-909.
Original article in IJSB Online
2 In Kilian's opinion [1] the change of the well-established name Pseudomonas betle may be a source of confusion. So, with reference to the first Principle of the Bacteriological Code (1990 Revision), Kilian requested that the original name Pseudomonas betle be conserved [2]. The Judicial Commission denied this request, and no Opinion will be issued upon this request [3].
References:
1 KILIAN (M.): Necessary changes of bacterial names? ASM News, 1998, 64, 670.
2 KILIAN (M.): Recommended conservation of the names Streptococcus sanguis, Streptococcus rattus, Streptococcus cricetus, and seven other names included in the Approved Lists of Bacterial Names. Request for an Opinion. Int. J. Syst. Evol. Microbiol. 2001, 51, 723-724.
Original article by Killian 2001 in IJSEM Online
3 DE VOS (P.), TRÜPER (H.G.) and TINDALL (B.J.): Judicial Commission of the International Committee on Systematics of Prokaryotes Xth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 28, 29 and 31 July and 1 August 2002, Paris, France. Int. J. Syst. Evol. Microbiol., 2005, 55, 525-532.
Original article by De Vos et al. 2005 in IJSEM Online
3 Pseudomonas beteli corrig. (Ragunathan 1928) Savulescu 1947 (Approved Lists 1980) and ¤ Pseudomonas hibiscicola Moniz 1963 (Approved Lists 1980) are misnamed and should be transferred to the genus ¤ Xanthomonas Dowson 1939 (Approved Lists 1980). Pseudomonas beteli corrig., Pseudomonas hibiscicola and ¤ Xanthomonas maltophilia (Hugh 1981) Swings et al. 1983 are heterotypic synonyms. According to the Bacteriological Code (1990 revision) Pseudomonas hibiscicola and Xanthomonas maltophilia should be renamed Xanthomonas beteli because beteli is the oldest specific epithet. The authors do not feel however the need to formally propose nomenclatural changes.
Reference: VAN DEN MOOTER (M.) and SWINGS (J.): Numerical analysis of 295 phenotypic features of 266 Xanthomonas strain and related strains and an improved taxonomy of the genus. Int. J. Syst. Bacteriol., 1990, 40, 348-369.
Original article in IJSEM Online

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Pseudomonas borbori Vanparys et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) R-20821 = DSM 17834 = LMG 23199.
Sequence accession no. (16S rRNA gene) for the type strain: AM114527.
Etymology: Gr. n. borboros, sludge; N.L. gen. n. borbori, of sludge.
Valid publication: VANPARYS (B.), HEYLEN (K.), LEBBE (L.) and DE VOS (P.): Pseudomonas peli sp. nov. and Pseudomonas borbori sp. nov., isolated from a nitrifying inoculum. Int. J. Syst. Evol. Microbiol., 2006, 56, 1875-1881.
Original article in IJSEM Online

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Pseudomonas boreopolis Gray and Thornton 1928, species.
Type strain: (see also StrainInfo.net) ATCC 33662= CCUG 3365 = CIP 106717 = JCM 13306 = LMG 979 = NCIB (now NCIMB) 9401.
Sequence accession no. (16S rRNA gene) for the type strain: AB021391.
Etymology: Gr. n. boreas, the north; Gr. n. polis, a city; N.L. gen. n. boreopolis, of North City.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: GRAY (P.H.H.) and THORNTON (H.G.): Soil bacteria that decompose certain aromatic compounds. Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung II, 1928, 73, 74-96.

Note: According to Anzai et al., Pseudomonas boreopolis Gray and Thornton 1928 (Approved Lists 1980) does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

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Pseudomonas brassicacearum Achouak et al. 2000, sp. nov.
Type strain: (see also StrainInfo.net) DBK11 = CFBP 11706 = CIP 107059 = DSM 13227 = JCM 11938.
Sequence accession no. (16S rRNA gene) for the type strain: AF100321.
Etymology: N.L. fem. pl. gen. n. brassicacearum, of the Brassicaceae, referring to its isolation from the rhizoplane of plants belonging to the Brassicaceae.
Valid publication: ACHOUAK (W.), SUTRA (L.), HEULIN (T.), MEYER (J.M.), FROMIN (N.), DEGRAEVE (S.), CHRISTEN (R.) and GARDAN (L.): Pseudomonas brassicacearum sp. nov. and Pseudomonas thivervalensis sp. nov., two root-associated bacteria isolated from Brassica napus and Arabidopsis thaliana. Int. J. Syst. Evol. Microbiol., 2000, 50, 9-18.
Original article in IJSEM Online

Pseudomonas brassicacearum Achouak et al. 2000 emend. Ivanova et al. 2009.
Type strain: (see also StrainInfo.net) DBK11 = CFBP 11706 = CIP 107059 = DSM 13227 = JCM 11938.
Sequence accession no. (16S rRNA gene) for the type strain: AF100321.
Etymology: N.L. fem. pl. gen. n. brassicacearum, of the Brassicaceae, referring to its isolation from the rhizoplane of plants belonging to the Brassicaceae.
Reference: IVANOVA (E.P.), CHRISTEN (R.), BIZET (C.), CLERMONT (D.), MOTREFF (L.), BOUCHIER (C.), ZHUKOVA (N.V.), CRAWFORD (R.J.) and KIPRIANOVA (E.A.): Pseudomonas brassicacearum subsp. neoaurantiaca subsp. nov., orange-pigmented bacteria isolated from soil and the rhizosphere of agricultural plants. Int. J. Syst. Evol. Microbiol., 2009, 59, 2476-2481.
Original article in IJSEM Online

Note: Subsequently, this species has been divided into subspecies (see: below).

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Pseudomonas brassicacearum subsp. brassicacearum (Achouak et al. 2000) Ivanova et al. 2009, subsp. nov.
Type strain: (see also StrainInfo.net) DBK11 = CFBP 11706 = CIP 107059 = DSM 13227 = JCM 11938.
Sequence accession no. (16S rRNA gene) for the type strain: AF100321.
Etymology: N.L. fem. pl. gen. n. brassicacearum, of the Brassicaceae, referring to its isolation from the rhizoplane of plants belonging to the Brassicaceae.
Valid publication: IVANOVA (E.P.), CHRISTEN (R.), BIZET (C.), CLERMONT (D.), MOTREFF (L.), BOUCHIER (C.), ZHUKOVA (N.V.), CRAWFORD (R.J.) and KIPRIANOVA (E.A.): Pseudomonas brassicacearum subsp. neoaurantiaca subsp. nov., orange-pigmented bacteria isolated from soil and the rhizosphere of agricultural plants. Int. J. Syst. Evol. Microbiol., 2009, 59, 2476-2481.
Original article in IJSEM Online

Note: The original authorship, Achouak et al. 2000, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online

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Pseudomonas brassicacearum subsp. neoaurantiaca Ivanova et al. 2009, subsp. nov.
Type strain: (see also StrainInfo.net) ATCC 49054= CIP 109457 = IMV 387 = VKM B-1524.
Sequence accession no. (16S rRNA gene) for the type strain: EU391388.
Etymology: Gr. adj. neos, new; N.L. fem. adj. aurantiaca, orange-coloured and also a bacterial specific epithet; N.L. fem. adj. neoaurantiaca, new aurantiaca, referring to the phenotypic similarity to Pseudomonas aurantiaca.
Valid publication: IVANOVA (E.P.), CHRISTEN (R.), BIZET (C.), CLERMONT (D.), MOTREFF (L.), BOUCHIER (C.), ZHUKOVA (N.V.), CRAWFORD (R.J.) and KIPRIANOVA (E.A.): Pseudomonas brassicacearum subsp. neoaurantiaca subsp. nov., orange-pigmented bacteria isolated from soil and the rhizosphere of agricultural plants. Int. J. Syst. Evol. Microbiol., 2009, 59, 2476-2481.
Original article in IJSEM Online

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Pseudomonas brenneri Baïda et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFML 97-391 = CIP 106646 = DSM 15294 = JCM 13307.
Sequence accession no. (16S rRNA gene) for the type strain: AF268968.
Etymology: N.L. masc. gen. n. brenneri, of Brenner, in honor of D.J. Brenner, an American microbiologist, for his contributions to the taxonomy of the families Enterobacteriaceae and Legionellaceae.
Valid publication: Validation List no. 87. Int. J. Syst. Evol. Microbiol., 2002, 52, 1437-1438.
Effective publication: BAÏDA (N.), YAZOURH (A.), SINGER (E.) and IZARD (D.): Pseudomonas brenneri sp. nov., a new species isolated from natural mineral waters. Res. Microbiol., 2001, 152, 493-502.
Validation List no. 87 in IJSEM Online - Effective publication Online

Note: According to Rules 27(3) and 30, this name is not validly published because the effective publication only documents deposit of the type strain in a single recognized culture collection. However, according to the Judicial Opinion 81, Pseudomonas brenneri Baïda et al. 2002 is to be considered to be validly published.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online

Pseudomonas brenneri Baïda et al. 2002 emend. van den Beld et al. 2016.
Reference: VAN DEN BELD, M. J., REINDERS, E., NOTERMANS, D. W. and REUBSAET, F. A. 2016. Possible misidentification of species in the Pseudomonas fluorescens lineage as Burkholderia pseudomallei and Francisella tularensis, and emended descriptions of Pseudomonas brenneri, Pseudomonas gessardii and Pseudomonas proteolytica. Int. J. Syst. Evol. Microbiol., 66, 3420-3425.
Original article.

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Pseudomonas caeni Xiao et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) HY-14 = CCTCC AB208156 = KCTC 22292.
Sequence accession no. (16S rRNA gene) for the type strain: EU620679.
Etymology: L. gen. neut. n. caeni, of sludge.
Valid publication: XIAO (Y.P.), HUI (W.), WANG (Q.), ROH (S.W.), SHI (X.Q.), SHI (J.H.) and QUAN (Z.X.): Pseudomonas caeni sp. nov., a denitrifying bacterium isolated from the sludge of an anaerobic ammonium-oxidizing bioreactor. Int. J. Syst. Evol. Microbiol., 2009, 59, 2594-2598.
Original article in IJSEM Online

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Pseudomonas cannabina (ex Šutič and Dowson 1959) Gardan et al. 1999, sp. nov., nom. rev.
Type strain: (see also StrainInfo.net) CFBP 2341 = CIP 106140 = ICMP 2823 = LMG 5096 = NCPPB 1437.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492827.
Synonym: "Pseudomonas cannabina" Šutič and Dowson 1959.
Etymology: L. fem. adj. cannabina, of hemp, of Cannabis, the generic name of the host plant, Cannabis sativa L.
Valid publication: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Sutic and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

Note: Pseudomonas cannabina Gardan et al. 1999 includes Pseudomonas syringae pv. Cannabina.

Pseudomonas cannabina (ex Šutič and Dowson 1959) Gardan et al. 1999 emend. Bull et al. 2010.
Type strain: (see also StrainInfo.net) CFBP 2341 = CIP 106140 = ICMP 2823 = LMG 5096 = NCPPB 1437.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492827.
Synonym: "Pseudomonas cannabina" Šutič and Dowson 1959.
Etymology: L. fem. adj. cannabina, of hemp, of Cannabis, the generic name of the host plant, Cannabis sativa L.
References: List of Changes in Taxonomic Opinion no. 12. Int. J. Syst. Evol. Microbiol., 2010, 60, 1482-1483 [2. BULL (C.T.), MANCEAU (C.), LYDON (J.), KONG (H.), VINATZER (B.A.) and FISCHER-LE SAUX (M.): Pseudomonas cannabina pv. cannabina pv. nov., and Pseudomonas cannabina pv. alisalensis (Cintas Koike and Bull, 2000) comb. nov., are members of the emended species Pseudomonas cannabina (ex Šutič & Dowson 1959) Gardan, Shafik, Belouin, Brosch, Grimont & Grimont 1999. Syst. Appl. Microbiol., 2010, 33, 105-115.]
List of Changes in Taxonomic Opinion no. 12 in IJSEM Online - Effective publication Online

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Pseudomonas carboxydohydrogena (ex Sanjieva and Zavarzin 1971) Meyer et al. 1980, nom. rev., comb. nov.
Type strain: (see also StrainInfo.net) "Carboxydobacterium" Z-1062 = ATCC 29978= DSM 1083 = JCM 12568 = VKM B-1334.
Sequence accession no. (16S rRNA gene) for the type strain: AB021393.
Synonym: "Seliberia carboxydohydrogena" Sanjieva and Zavarzin 1971.
Etymology: not given in the paper by Meyer et al. 1980.
Valid publication: MEYER (O.), LALUCAT (J.) and SCHLEGEL (H.G.): Pseudomonas carboxydohydrogena (Sanjieva and Zavarzin) comb. nov., a monotrichous, nonbudding, strictly aerobic, carbon monoxide-utilizing hydrogen bacterium previously assigned to Seliberia. Int. J. Syst. Bacteriol., 1980, 30, 189-195.
Original article in IJSEM Online

Note: According to Anzai et al., Pseudomonas carboxydohydrogena (ex Sanjieva and Zavarzin 1971) Meyer et al. 1980 does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

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Pseudomonas caricapapayae Robbs 1956, species.
Type strain: (see also StrainInfo.net) ATCC 33615= CCUG 32775 = CFBP 3204 = CIP 106736 = ICMP 2855 = LMG 2152 = NCPPB 1873.
Sequence accession no. (16S rRNA gene) for the type strain: D84010.
Etymology: N.L. gen. n. caricapapayae, of Carica papaya, pawpaw.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: ROBBS (C.F.): Uma nova doenca bacteriana do mamoeiro. Revista da Sociedade Brasileira de Agronomia, 1956, 12, 73-76.

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Pseudomonas caryophylli (Burkholder 1942) Starr and Burkholder 1942, species.
Type strain: (see also StrainInfo.net) ATCC 25418= CCUG 20834 = CFBP 2429 = CFBP 3818 = CIP 105770 = DSM 50341 = HAMBI 2159 = ICMP 512 = JCM 9310 = JCM 10488 = LMG 2155 = NCPPB 2151.
Sequence accession no. (16S rRNA gene) for the type strain: AB021423.
Synonym: "Phytomonas caryophylli" Burkholder 1942.
Etymology: N.L. masc. n. caryophyllus, specific epithet of Dianthus caryophyllus, carnation; N.L. gen. n. caryophylli, of the carnation.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 BURKHOLDER (W.H.): Three bacterial plant pathogens, Phytomonas caryophylli, sp. n., Phytomonas alliicola sp. n., and Phytomonas manihoti (Arthaud-Berthet et Bondar) Viegas. Phytopathology, 1942, 32, 141-149. 2 STARR (M.P.) and BURKHOLDER (W.H.): Lipolytic activity of phytopathogenic bacteria determined by means of spirit blue agar and its taxonomic significance. Phytopathology, 1942, 32, 598-604.

¤ Burkholderia caryophylli (Burkholder 1942) Yabuuchi et al. 1993, comb. nov.

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Pseudomonas cattleyae (Pavarino 1911) Savulescu 1947, species.
Type strain: (see also StrainInfo.net) FC-507 = ATCC 33619= CCUG 21975 = CFBP 2423 = CIP 106435 = IBSBF 209 = ICMP 2826 = ICPB 30134 (PC21) = LMG 2364 = NCPPB 961.
Sequence accession no. (16S rRNA gene) for the type strain: AF137504.
Synonym: "Bacterium cattleyae" Pavarino 1911.
Etymology: N.L. n. Cattleya, a genus of orchids; N.L. gen. n. cattleyae, of Cattleya.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 PAVARINO (G.L.): Malattie causate da bacteri nelle orchidee. Atti della Reale Accademia dei Lincei, 1911, 20, 233-237. 2 SAVULESCU (T.): Contribution à la classification des bacteriacées phytopathogènes. Analele Academiei Romane Series III Tom 22, Memoire, 1947, 4, 1-26.

¤ Pseudomonas avenae subsp. avenae Manns 1909 (Approved Lists 1980).

Note: Because the type strain of Pseudomonas cattleyae (Pavarino 1911) Savulescu 1947 (Approved Lists 1980) is allocated to Pseudomonas avenae subsp. avenae Manns 1909 (Approved Lists 1980), Hu et al. 1991 propose Pseudomonas cattleyae (Pavarino 1911) Savulescu 1947 (Approved Lists 1980) as a later heterotypic synonym of Pseudomonas avenae Manns 1909 (Approved Lists 1980).
Reference: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

¤ Acidovorax avenae subsp. cattleyae (Pavarino 1911) Willems et al. 1992, comb. nov.

¤ Acidovorax cattleyae (Pavarino 1911) Schaad et al. 2009, comb. nov.

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Pseudomonas cedrina corrig. Dabboussi et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFML 96-198 = CIP 105541 = DSM 17516.
Sequence accession no. (16S rRNA gene) for the type strain: AF064461.
Etymology: L. fem. adj. cedrina, of the cedar tree, the characteristic tree of Lebanon, the country from which isolates were taken.
Valid publication: Validation List no. 87. Int. J. Syst. Evol. Microbiol., 2002, 52, 1437-1438.
Effective publication: DABBOUSSI (F.), HAMZE (M.), ELOMARI (M.), VERHILLE (S.), BAIDA (N.), IZARD (D.) and LECLERC (H.): Taxonomic study of bacteria isolated from Lebanese spring waters: proposal of Pseudomonas cedrella sp. nov. and P. orientalis sp. nov. Res. Microbiol., 1999, 150, 303-316.
Validation List no. 87 in IJSEM Online - Effective publication Online

Notes:
1 The original spelling of the specific epithet, cedrella (sic), has been corrected on validation according to Rule 61.
Reference: ASSOCIATE EDITOR, IJSEM: Validation List no. 87 (footnote §). Int. J. Syst. Evol. Microbiol., 2002, 52, 1437-1438.
2 At the time of publication, the type strain was not deposited in two publicly accessible service collections in different countries, and Pseudomonas cedrina corrig. Dabboussi et al. 2002 was not validly published [Rules 27(3) and 30]. However, according to the Judicial Opinion 81, Pseudomonas cedrina corrig. Dabboussi et al. 2002 is now considered to be validly published, since deposit in a second culture collection in a different country has been documented.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online
3 Subsequently, this species has been divided into subspecies (see: below).

Pseudomonas cedrina corrig. Dabboussi et al. 2002 emend. Behrendt et al. 2009.
Type strain: (see also StrainInfo.net) CFML 96-198 = CIP 105541 = DSM 17516.
Sequence accession no. (16S rRNA gene) for the type strain: AF064461.
Etymology: L. fem. adj. cedrina, of the cedar tree, the characteristic tree of Lebanon, the country from which isolates were taken.
Reference: BEHRENDT (U.), SCHUMANN (P.), MEYER (J.M.) and ULRICH (A.): Pseudomonas cedrina subsp. fulgida subsp. nov., a fluorescent bacterium isolated from the phyllosphere of grasses; emended description of Pseudomonas cedrina and description of Pseudomonas cedrina subsp. cedrina subsp. nov. Int. J. Syst. Evol. Microbiol., 2009, 59, 1331-1335.
Original article in IJSEM Online

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Pseudomonas cedrina subsp. cedrina (Dabboussi et al. 2002) Behrendt et al. 2009, subsp. nov.
Type strain: (see also StrainInfo.net) CFML 96-198 = CIP 105541 = DSM 17516.
Sequence accession no. (16S rRNA gene) for the type strain: AF064461.
Etymology: L. fem. adj. cedrina, of the cedar tree, the characteristic tree of Lebanon, the country from which isolates were taken.
Valid publication: BEHRENDT (U.), SCHUMANN (P.), MEYER (J.M.) and ULRICH (A.): Pseudomonas cedrina subsp. fulgida subsp. nov., a fluorescent bacterium isolated from the phyllosphere of grasses; emended description of Pseudomonas cedrina and description of Pseudomonas cedrina subsp. cedrina subsp. nov. Int. J. Syst. Evol. Microbiol., 2009, 59, 1331-1335.
Original article in IJSEM Online

Note: The original authorship, Dabboussi et al. 2002, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online

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Pseudomonas cedrina subsp. fulgida Behrendt et al. 2009, subsp. nov.
Type strain: (see also StrainInfo.net) P 515/12 = DSM 14938 = LMG 21467.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492830.
Etymology: L. fem. adj. fulgida, flashing, glittering, shining, pertaining to the bright fluorescence.
Valid publication: BEHRENDT (U.), SCHUMANN (P.), MEYER (J.M.) and ULRICH (A.): Pseudomonas cedrina subsp. fulgida subsp. nov., a fluorescent bacterium isolated from the phyllosphere of grasses; emended description of Pseudomonas cedrina and description of Pseudomonas cedrina subsp. cedrina subsp. nov. Int. J. Syst. Evol. Microbiol., 2009, 59, 1331-1335.
Original article in IJSEM Online

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Pseudomonas cepacia (ex Burkholder 1950) Palleroni and Holmes 1981, sp. nov., nom. rev.
Type strain: (see also StrainInfo.net) Ballard 717 = 717-ICPB 25 = ATCC 25416= CCUG 12691 = CCUG 13226 = CFBP 2227 = CIP 80.24 = DSM 7288 = HAMBI 1976 = ICMP 5796 = IFO (now NBRC) 14074 = JCM 5964 = LMG 1222 = NCCB 76047 = NCPPB 2993 = NCTC 10743 = NRRL B-14810.
Sequence accession no. (16S rRNA gene) for the type strain: U96927.
Synonyms: "Pseudomonas cepacia" Burkholder 1950, "Pseudomonas multivorans" Stanier et al. 1966.
Etymology: L. fem. n. caepa or cepa, onion; N.L. fem. adj. cepacia, of or like an onion.
Valid publication: PALLERONI (N.J.) and HOLMES (B.): Pseudomonas cepacia sp. nov., nom. rev. Int. J. Syst. Bacteriol., 1981, 31, 479-481.
Original article in IJSEM Online

¤ Burkholderia cepacia (Palleroni and Holmes 1981) Yabuuchi et al. 1993, comb. nov.

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Pseudomonas cerasi Kaluzna et al. 2017, sp. nov.
Type strain: (see also StrainInfo.net) 58=CFBP 8305=LMG 28609.
Sequence accession no. (16S rRNA) for the type strain: LN713316.
Whole-genome sequence accession no.: NZ_LT222319.
Etymology: N/A.
Source: Plant pathogen.
Valid publication: VALIDATION LIST no. 173. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 67 (2017), 1-3.
Validation List Online
Effective publication: KALUZNA, M., WILLEMS, A., POTHIER, J. F., RUINELLI, M., SOBICZEWSKI, P. and PULAWSKA, J. 2016. Pseudomonas cerasi sp. nov. (non Griffin, 1911) isolated from diseased tissue of cherry. Syst. Appl. Microbiol., 39, 370-377.
Effective publication Online

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Pseudomonas chengduensis Tao et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) MBR=CGMCC 2318=DSM 26382.
Sequence accession no. (16S rRNA) for the type strain: EU307111.
Etymology: N.L. fem. adj. chengduensis, pertaining to Chengdu, where the type strain was isolated.
Source: Environmental.
Valid publication: TAO, Y., ZHOU, Y., HE, X., HU, X. and LI, D. 2014. Pseudomonas chengduensis sp. nov., isolated from landfill leachate. Int. J. Syst. Evol. Microbiol., 64, 95-100.
Original article in IJSEM Online

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Pseudomonas chloritidismutans Wolterink et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) AW-1 = ATCC BAA-443 = DSM 13592 = NBRC 102227.
Sequence accession no. (16S rRNA gene) for the type strain: AY017341.
Etymology: N.L. n. chloris, chlorite; N.L. part. adj. dismutans, splitting; N.L. part. adj. chloritidismutans, chlorite-splitting.
Valid publication: WOLTERINK (A.F.W.M.), JONKER (A.B.), KENGEN (S.W.M.) and STAMS (A.J.M.): Pseudomonas chloritidismutans sp. nov., a non-denitrifying, chlorate-reducing bacterium. Int. J. Syst. Evol. Microbiol., 2002, 52, 2183-2190.
Original article in IJSEM Online

¤ Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980).

Note: According to Cladera et al. 2006, Pseudomonas chloritidismutans Wolterink et al. 2002 is a later heterotypic synonym of ¤ Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980).
References: List of Changes in Taxonomic Opinion no. 4. Int. J. Syst. Evol. Microbiol., 2006, 56, 1463. [CLADERA (A.M.), GARCÍA-VALDÉS (E.) and LALUCAT (J.): Genotype versus phenotype in the circumscription of bacterial species: the case of Pseudomonas stutzeri and Pseudomonas chloritidismutans. Arch. Microbiol., 2006, 184, 353-361.]
List of Changes in Taxonomic Opinion no. 4 in IJSEM Online - Effective publication Online

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Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930, species.
Type strain: (see also StrainInfo.net) ATCC 9446= CCUG 552 B = CFBP 2132 = CIP 63.22 = DSM 50083 = HAMBI 2011 = IFO (now NBRC) 3904 = JCM 2778 = LMG 5004 = NCCB 76041 = NCIMB 9392 = NCTC 13002 = VKM B-1246.
Sequence accession no. (16S rRNA gene) for the type strain: Z76673.
Synonym: "Bacillus chlororaphis" Guignard and Sauvageau 1894.
Etymology: Gr. adj. chloros, green; Gr. n. raphis, a needle; N.L. n. chlororaphis, a green needle.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 GUIGNARD (L.) and SAUVAGEAU (C.): Sur un nouveau microbe chromogène, le Bacillus chloroaphis. Comptes Rendus des Séances de la Société de Biologie (Paris), Séries 10, 1894, 1, 841-843. 2 BERGEY (D.H.), HARRISON (F.C.), BREED (R.S.), HAMMER (B.W.) and HUNTOON (F.M.) (editors): Bergey's Manual of Determinative Bacteriology, 3rd ed., The Williams & Wilkins Co, Baltimore, 1930, pp. 1-589.

Notes:
1 According to Johnson and Palleroni 1989, Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas aureofaciens Kluyver 1956 (Approved Lists 1980).
Reference: JOHNSON (J.L.) and PALLERONI (N.J.): Deoxyribonucleic acid similarities among Pseudomonas species. Int. J. Syst. Bacteriol., 1989, 39, 230-235.
Original article in IJSEM Online
2 Subsequently, this species has been divided into subspecies (see: below).

Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930 (Approved Lists 1980) emend. Peix et al. 2007.
Type strain: (see also StrainInfo.net)StrainInfo.net) ATCC 9446= CCUG 552 B = CFBP 2132 = CIP 63.22 = DSM 50083 = HAMBI 2011 = IFO (now NBRC) 3904 = JCM 2778 = LMG 5004 = NCCB 76041 = NCIMB 9392 = NCTC 13002 = VKM B-1246.
Sequence accession no. (16S rRNA gene) for the type strain: Z76673.
Synonym: "Bacillus chlororaphis" Guignard and Sauvageau 1894.
Etymology: Gr. adj. chloros, green; Gr. n. raphis, a needle; N.L. n. chlororaphis, a green needle.
Reference: PEIX (A.), VALVERDE (A.), RIVAS (R.), IGUAL (J.M.), RAMÍREZ-BAHENA (M.H.), MATEOS (P.F.), SANTA-REGINA (I.), RODRÍGUEZ-BARRUECO (C.), MARTÍNEZ-MOLINA (E.) and VELÁZQUEZ (E.): Reclassification of Pseudomonas aurantiaca as a synonym of Pseudomonas chlororaphis and proposal of three subspecies, P. chlororaphis subsp. chlororaphis subsp. nov., P. chlororaphis subsp. aureofaciens subsp. nov., comb. nov. and P. chlororaphis subsp. aurantiaca subsp. nov., comb. nov. Int. J. Syst. Evol. Microbiol., 2007, 57, 1286-1290.
Original article in IJSEM Online

Note: In the paper by Peix et al. 2007, Pseudomonas chlororaphis (Guignard and Sauvageau 1894) Bergey et al. 1930 is cited as Pseudomonas chlororaphis (ex Guignard and Sauvageau 1894) Bergey et al. 1930.

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Pseudomonas chlororaphis subsp. aurantiaca (Nakhimovskaya 1948) Peix et al. 2007, comb. nov.
Type strain: (see also StrainInfo.net) ATCC 33663= CIP 106718 = NCIMB 10068 = VKM B-876.
Sequence accession no. (16S rRNA gene) for the type strain: DQ682655.
Basonym: ¤ Pseudomonas aurantiaca Nakhimovskaya 1948 (Approved Lists 1980).
Etymology: N.L. fem. adj. aurantiaca, orange coloured.
Valid publication: PEIX (A.), VALVERDE (A.), RIVAS (R.), IGUAL (J.M.), RAMÍREZ-BAHENA (M.H.), MATEOS (P.F.), SANTA-REGINA (I.), RODRÍGUEZ-BARRUECO (C.), MARTÍNEZ-MOLINA (E.) and VELÁZQUEZ (E.): Reclassification of Pseudomonas aurantiaca as a synonym of Pseudomonas chlororaphis and proposal of three subspecies, P. chlororaphis subsp. chlororaphis subsp. nov., P. chlororaphis subsp. aureofaciens subsp. nov., comb. nov. and P. chlororaphis subsp. aurantiaca subsp. nov., comb. nov. Int. J. Syst. Evol. Microbiol., 2007, 57, 1286-1290.
Original article in IJSEM Online

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Pseudomonas chlororaphis subsp. aureofaciens (Kluyver 1956) Peix et al. 2007, comb. nov.
Type strain: (see also StrainInfo.net)StrainInfo.net) ATCC 13985= CFBP 2133 = DSM 6698 = LMG 1245 = NCIMB 9030.
Sequence accession no. (16S rRNA gene) for the type strain: AY509898.
Basonym: ¤ Pseudomonas aureofaciens Kluyver 1956 (Approved Lists 1980).
Etymology: L. adj. aureus, golden; L. part. adj. faciens, making; N.L. part. adj. aureofaciens, making golden.
Valid publication: PEIX (A.), VALVERDE (A.), RIVAS (R.), IGUAL (J.M.), RAMÍREZ-BAHENA (M.H.), MATEOS (P.F.), SANTA-REGINA (I.), RODRÍGUEZ-BARRUECO (C.), MARTÍNEZ-MOLINA (E.) and VELÁZQUEZ (E.): Reclassification of Pseudomonas aurantiaca as a synonym of Pseudomonas chlororaphis and proposal of three subspecies, P. chlororaphis subsp. chlororaphis subsp. nov., P. chlororaphis subsp. aureofaciens subsp. nov., comb. nov. and P. chlororaphis subsp. aurantiaca subsp. nov., comb. nov. Int. J. Syst. Evol. Microbiol., 2007, 57, 1286-1290.
Original article in IJSEM Online

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Pseudomonas chlororaphis subsp. chlororaphis (Guignard and Sauvageau 1894) Peix et al. 2007, subsp. nov.
Type strain: (see also StrainInfo.net) ATCC 9446= CCUG 552 B = CFBP 2132 = CIP 63.22 = DSM 50083 = HAMBI 2011 = IFO (now NBRC) 3904 = JCM 2778 = LMG 5004 = NCCB 76041 = NCIMB 9392 = NCTC 13002 = VKM B-1246.
Sequence accession no. (16S rRNA gene) for the type strain: Z76673.
Etymology: Gr. adj. chloros, green; Gr. n. raphis, a needle; N.L. n. chlororaphis, a green needle.
Valid publication: PEIX (A.), VALVERDE (A.), RIVAS (R.), IGUAL (J.M.), RAMÍREZ-BAHENA (M.H.), MATEOS (P.F.), SANTA-REGINA (I.), RODRÍGUEZ-BARRUECO (C.), MARTÍNEZ-MOLINA (E.) and VELÁZQUEZ (E.): Reclassification of Pseudomonas aurantiaca as a synonym of Pseudomonas chlororaphis and proposal of three subspecies, P. chlororaphis subsp. chlororaphis subsp. nov., P. chlororaphis subsp. aureofaciens subsp. nov., comb. nov. and P. chlororaphis subsp. aurantiaca subsp. nov., comb. nov. Int. J. Syst. Evol. Microbiol., 2007, 57, 1286-1290.
Original article in IJSEM Online

Note: The original authorship, (Guignard and Sauvageau 1894) Bergey et al. 1930, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online

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Pseudomonas chlororaphis subsp. piscium Burr et al. 2010, subsp. nov.
Type strain: (see also StrainInfo.net) JF3835 = DSM 21509 = NCIMB 14478.
Sequence accession no. (16S rRNA gene) for the type strain: FJ168539.
Etymology: L. n. piscis, fish; L. gen. pl. n. piscium, of/from fishes.
Valid publication: BURR (S.E.), GOBELI (S.), KUHNERT (P.), GOLDSCHMIDT-CLERMONT (E.), and FREY (J.): Pseudomonas chlororaphis subsp. piscium subsp. nov., isolated from freshwater fish. Int. J. Syst. Evol. Microbiol., 2010, 60, 2753-2757.
Original article in IJSEM Online

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Pseudomonas cichorii (Swingle 1925) Stapp 1928, species.
Type strain: (see also StrainInfo.net) ATCC 10857= CCUG 32776 = CFBP 2101 = CIP 106704 = DSM 50259 = ICMP 5707 = LMG 2162 = NCCB 69031 = NCPPB 943.
Sequence accession no. (16S rRNA gene) for the type strain: Z76658.
Synonyms: "Phytomonas cichorii" Swingle 1925, "Bacterium cichorii" (Swingle 1925) Elliott 1930, "Pseudomonas endiviae" Kotte 1930, "Bacterium formosanum" Okabe 1935, "Chlorobacter cichorii" (Swingle 1925) Patel and Kulkarni 1951, probable synonym: "Pseudomonas papaveris" Lelliott and Wallace 1955.
Etymology: L. n. cichorium, chicory, succory, or endive; L. gen. n. cichorii, of chicory.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SWINGLE (D.B.): Center rot of "french endive" or wilt of chicory (Cichorium intybos L.). Phytopathology, 1925, 15, 730 (Abstr.). 2 STAPP (C.): Schizomycetes (Spaltpilze oder Bakterien). In: SORAUER (ed.), Handbuch der Pflanzenkrankheiten, 5th ed., Vol. 2, Paul Parey, Berlin, 1928, pp. 1-295.

Note: On the Approved Lists of Bacterial Names and on the Approved Lists of Bacterial Names (Amended Edition), the page numbers of the paper by Stapp are cited as pp. 758.

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Pseudomonas cissicola (Takimoto 1939) Burkholder 1948, species.
Type strain: (see also StrainInfo.net) ATCC 33616= CCUG 18839 = CFBP 2432 = CIP 106723 = JCM 13362 = NCPPB 2982.
Sequence accession no. (16S rRNA gene) for the type strain: AB021399.
Synonym: "Aplanobacter cissicola" Takimoto 1939.
Etymology: N.L. n. Cissus, generic name of flowering plant; L. suff. -cola (from L. n. incola), dweller; N.L. n. cissicola, Cissus dweller.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 TAKIMOTO (S.): Bacterial leaf spot of Cissus japonica Willd. Annals of the Phytopathological Society, Japan, 1939, 9, 41-43. 2 BURKHOLDER (W.H.): Genus I. Pseudomonas Migula. Bacterial plant pathogens. In: R.S. BREED, E.G.D. MURRAY and A.P. HITCHENS (editors): Bergey's Manual of Determinative Bacteriology, 6th ed., The Williams & Wilkins Co, Baltimore, 1948, pp. 82-150.

Notes:
1 The type strain was obviously mislabeled at several culture collections. These errors have caused considerable confusion in the taxonomic placement of Pseudomonas cissicola. The type strain of Pseudomonas cissicola should be strain PC1 in the culture collection of plant-pathogenic bacteria, Laboratory of Plant Pathology, Shizuoka University (836 Ohya, Shizuoka 422, Japan) [1], strain ATCC 33616 [1], strain CCM 2888 [1], strain CFBP 2432 [2], ICMP 8561 [2], LMG 2167 [2], or NCPPB 2982 [2].
References:
1 GOTO (M.): Taxonomic confusion of Pseudomonas cissicola originated from mislabeling by International Type Culture Collections. Int. J. Syst. Bacteriol., 1992, 42, 503-505.
Original article in IJSEM Online
2 HU (F.P.), YOUNG (J.M.), STEAD (D.E.) and GOTO (M.): Transfer of Pseudomonas cissicola (Takimoto 1939) Burkholder 1948 to the genus Xanthomonas. Int. J. Syst. Bacteriol., 1997, 47, 228-230.
Original article in IJSB Online
2 The type strain of Pseudomonas cissicola (Takimoto 1939) Burkholder 1948 has the determinative characteristics and the fatty acid profile of the genus Xanthomonas Dowson 1939 (Approved Lists 1980). However, it is not appropriate to propose the new combination Xanthomonas cissicola without investigating more extensively the genomic and phenotypic characteristics.
Reference: HU (F.P.), YOUNG (J.M.), STEAD (D.E.) and GOTO (M.): Transfer of Pseudomonas cissicola (Takimoto 1939) Burkholder 1948 to the genus Xanthomonas. Int. J. Syst. Bacteriol., 1997, 47, 228-230.
Original article in IJSB Online

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Pseudomonas citronellolis Seubert 1960, species.
Type strain: (see also StrainInfo.net) ATCC 13674= CCUG 17933 = CFBP 5585 = CIP 104381 = DSM 50332 = IAM 15129 = JCM 21587 = LMG 18378 = NBRC 103043 = NRRL B-2504.
Sequence accession no. (16S rRNA gene) for the type strain: Z76659.
Etymology: N.L. gen. n. citronellolis, of citronellol.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: SEUBERT (W.): Degradation of isoprenoid compounds by microorganisms. I. Isolation and characterization of an isoprenoid-degrading bacterium, Pseudomonas citronellolis n. sp. Journal of Bacteriology, 1960, 79, 426-434.

Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) emend. Lang et al. 2007.
Type strain: (see also StrainInfo.net) ATCC 13674= CCUG 17933 = CFBP 5585 = CIP 104381 = DSM 50332 = IAM 15129 = JCM 21587 = LMG 18378 = NRRL B-2504.
Sequence accession no. (16S rRNA gene) for the type strain: Z76659.
Etymology: N.L. gen. n. citronellolis, of citronellol.
Reference: LANG (E.), GRIESE (B.), SPRÖER (C.), SCHUMANN (P.), STEFFEN (M.) and VERBARG (S.): Characterization of 'Pseudomonas azelaica' DSM 9128, leading to emended descriptions of Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) and Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980), including Pseudomonas multiresinivorans as its later heterotypic synonym. Int. J. Syst. Evol. Microbiol., 2007, 57, 878-882.
Original article in IJSEM Online

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Pseudomonas cocovenenans van Damme et al. 1960, species.
Type strain: (see also StrainInfo.net) ATCC 33664= CFBP 4790 = DSM 11318 = JCM 10561 = LMG 11626 = NCIMB 9450.
Sequence accession no. (16S rRNA gene) for the type strain: AB021389.
Etymology: N.L. n. Cocos, genus of coconut; L. v. veneno, to poison; N.L. part. adj. cocovenenans, coconut poisoning.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: VAN DAMME (P.A.), JOHANNES (A.G.), COX (H.C.) and BERENDS (W.): On toxoflavin, the yellow poison of Pseudomonas cocovenenans. Recueil des Travaux Chimiques des Pays-Bas, 1960, 79, 255-267.

¤ Burkholderia cocovenenans (van Damme et al. 1960) Gillis et al. 1995, comb. nov.

¤ Burkholderia gladioli (Severini 1913) Yabuuchi et al. 1993.

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Pseudomonas coleopterorum Menéndez et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) Esc2Am=LMG 28558=CECT 8695.
Sequence accession no. (16S rRNA) for the type strain: KM888184.
Etymology: N.L. gen. pl. n. coleopterorum, of Coleoptera, referring to the order of insects from which the organism was first isolated.
Source: Animal.
Valid publication: MENÉNDEZ, E., RAMÍREZ-BAHENA, M. H., FABRYOVA, A., IGUAL, J. M., BENADA, O., MATEOS, P. F., PEIX, A., KOLAŘÍK, M. and GARCÍA-FRAILE, P. 2015. Pseudomonas coleopterorum sp. nov., a cellulase-producing bacterium isolated from the bark beetle Hylesinus fraxini. Int. J. Syst. Evol. Microbiol., 65, 2852-2858.
Original article in IJSEM Online

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Pseudomonas composti Gibello et al. 2011, sp. nov.
Type strain: (see also StrainInfo.net) C2 = CCUG 59231 = CECT 7516.
Sequence accession no. (16S rRNA gene) for the type strain: FN429930.
Etymology: N.L. n. compostum -i, compost; N.L. gen. n. composti, of compost, from which strains were first isolated.
Valid publication: GIBELLO (A.), VELA (A.I.), MARTÍN (M.), MENGS (G.), ALONSO (P.Z.), GARBI (C.) and FERNÁNDEZ-GARAYZÁBAL (J.F.): Pseudomonas composti sp. nov., isolated from compost samples. Int. J. Syst. Evol. Microbiol., 2011, 61, 2962-2966.
Original article in IJSEM Online

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Pseudomonas congelans Behrendt et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) P 538/23 = DSM 14939 = LMG 21466.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492828.
Etymology: L. part. adj. congelans, freezing/forming ice.
Valid publication: BEHRENDT (U.), ULRICH (A.) and SCHUMANN (P.): Fluorescent pseudomonads associated with the phyllosphere of grasses; Pseudomonas trivialis sp. nov., Pseudomonas poae sp. nov. and Pseudomonas congelans sp. nov. Int. J. Syst. Evol. Microbiol., 2003, 53, 1461-1469.
Original article in IJSEM Online

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Pseudomonas corrugata Roberts and Scarlett 1981, sp. nov.
Type strain: (see also StrainInfo.net) ATCC 29736= CCUG 23367 = CCUG 32778 = CFBP 2431 = CIP 105514 = DSM 7228 = ICMP 5819 = LMG 2172 = NCPPB 2445.
Sequence accession no. (16S rRNA gene) for the type strain: D84012.
Etymology: L. v. corrugare, to make full of wrinkles, to wrinkle up; L. fem. part. adj. corrugata, wrinkled up.
Valid publication: Validation List no. 6. Int. J. Syst. Bacteriol., 1981, 31, 215-218.
Effective publication: SCARLETT (C.M.), FLETCHER (J.T.), ROBERTS (P.) and LELLIOTT (R.A.): Tomato pith necrosis caused by Pseudomonas corrugata n. sp. Ann. Appl. Biol., 1978, 88, 105-114.
Validation List no. 6 in IJSEM Online

Pseudomonas corrugata Roberts and Scarlett 1981 emend. Sutra et al. 1997.
Type strain: (see also StrainInfo.net) ATCC 29736= CCUG 32778 = CFBP 2431 = CIP 105514 = DSM 7228 = ICMP 5819 = LMG 2172 = NCPPB 2445.
Sequence accession no. (16S rRNA gene) for the type strain: D84012.
Etymology: L. v. corrugare, to make full of wrinkles, to wrinkle up; L. fem. part. adj. corrugata, wrinkled up.
Reference: SUTRA (L.), SIVERIO (F.), LOPEZ (M.M.), HUNAULT (G.), BOLLET (C.) and GARDAN (L.): Taxonomy of Pseudomonas strains isolated from tomato pith necrosis: emended description of Pseudomonas corrugata and proposal of three unnamed fluorescent Pseudomonas genomospecies. Int. J. Syst. Bacteriol., 1997, 47, 1020-1033.
Original article in IJSEM Online

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Pseudomonas costantinii Munsch et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) PS 3a = CFBP 5705 = HAMBI 2444.
Sequence accession no. (16S rRNA gene) for the type strain: AF374472.
Etymology: N.L. gen. masc. n. costantinii, of Costantin, referring to Julien Costantin, a French mycologist who, in a note dated March 1892 to the "Société de Biologie", described some superficial brown blotch areas on sporophores.
Valid publication: MUNSCH (P.), ALATOSSAVA (T.), MARTTINEN (N.), MEYER (J.M.), CHRISTEN (R.) and GARDAN (L.): Pseudomonas costantinii sp. nov., another causal agent of brown blotch disease, isolated from cultivated mushroom sporophores in Finland. Int. J. Syst. Evol. Microbiol., 2002, 52, 1973-1983.
Original article in IJSEM Online

Note: The specific epithet, costantinii, is a N.L. gen. n. not a N.L. adj. as cited by Munsch et al. 2002.

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Pseudomonas cremoricolorata Uchino et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CC-8 = NRIC 0181 = CIP 107616 = DSM 17059 = IFO (now NBRC) 16634 = JCM 11246.
Sequence accession no. (16S rRNA gene) for the type strain: AB060137.
Etymology: L. n. cremor, the thick juice obtained from animal or vegetable substances; L. adj. coloratus -a -um, colored; N.L. fem. adj. cremoricolorata, intended to mean cream-colored.
Valid publication: Validation List no. 85. Int. J. Syst. Evol. Microbiol., 2002, 52, 685-690.
Effective publication: UCHINO (M.), SHIDA (O.), UCHIMURA (T.) and KOMAGATA (K.): Recharacterization of Pseudomonas fulva Iizuka and Komagata 1963, and proposals of Pseudomonas parafulva sp. nov. and Pseudomonas cremoricolorata sp. nov. J. Gen. Appl. Microbiol., 2001, 47, 247-261.
Validation List no. 85 in IJSEM Online - Effective publication Online

Notes:
1 In Validation List no. 85, the reference of the effective publication is erroneously cited as J. Gen. Appl. Microbiol., 2001, 46, 247-261.
2 According to Rules 27(3) and 30, this name is not validly published, because the original publication makes reference to the deposit of the type strain in a single recognized culture collection. However, according to the Judicial Opinion 81, Pseudomonas cremoricolorata Uchino et al. 2002 is considered to be validly published, since deposit in a second culture collection in a different country has been documented.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online

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Pseudomonas cuatrocienegasensis Escalante et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) 1N = CIP 109853 = LMG 24676.
Sequence accession no. (16S rRNA gene) for the type strain: EU791281.
Etymology: N.L. fem. adj. cuatrocienegasensis, pertaining to the Cuatro Ciénegas valley, the location of the evaporative lagoon from which the organism was isolated.
Valid publication: ESCALANTE (A.E.), CABALLERO-MELLADO (J.), MARTÍNEZ-AGUILAR (L.), RODRÍGUEZ-VERDUGO (A.), GONZÁLEZ-GONZÁLEZ (A.), TORIBIO-JIMÉNEZ (J.) and SOUZA (V.): Pseudomonas cuatrocienegasensis sp. nov., isolated from an evaporating lagoon in the Cuatro Cienegas valley in Coahuila, Mexico. Int. J. Syst. Evol. Microbiol., 2009, 59, 1416-1420.
Original article in IJSEM Online

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Pseudomonas deceptionensis Carrión et al. 2011, sp. nov.
Type strain: (see also StrainInfo.net) M1 = M1(2010b) = CECT 7677 = LMG 25555.
Sequence accession no. (16S rRNA gene) for the type strain: GU936597.
Etymology: N.L. fem. adj. deceptionensis, of or pertaining to Deception Island, Antarctic Sea.
Valid publication: CARRIÓN (O.), MIÑANA-GALBIS (D.), MONTES (M.J.) and MERCADÉ (E.): Pseudomonas deceptionensis sp. nov., a psychrotolerant bacterium from the Antarctic. Int. J. Syst. Evol. Microbiol., 2011, 61, 2401-2405.
Original article in IJSEM Online

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Pseudomonas delafieldii Davis 1970, species.
Type strain: (see also StrainInfo.net) ATCC 17505= CCUG 1779 = CFBP 2442 = CFBP 3609 = CIP 103303 = DSM 64 = LMG 5943.
Sequence accession no. (16S rRNA gene) for the type strain: AF078764.
Etymology: N.L. gen. masc. n. delafieldii, of Delafield, named after F.P. Delafield, who first isolated this organism.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DAVIS (D.H.). In: DAVIS (D.H.), STANIER (R.Y.), DOUDOROFF (M.) and. MANDEL (M.): Taxonomic studies on some Gram-negative polarly flagellated "hydrogen bacteria" and related species. Archiv fur Mikrobiologie, 1970, 70, 1-13.

¤ Acidovorax delafieldii (Davis 1970) Willems et al. 1990, comb. nov.

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Pseudomonas delhiensis Prakash et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) RLD-1 = CCM 7361 = MTCC 7601.
Sequence accession no. (16S rRNA gene) for the type strain: DQ339153.
Etymology: N.L. fem. adj. delhiensis, pertaining to Delhi, the place of isolation of the type strain.
Valid publication: PRAKASH (O.), KUMARI (K.) and LAL (R.): Pseudomonas delhiensis sp. nov., from a fly ash dumping site of a thermal power plant. Int. J. Syst. Evol. Microbiol., 2007, 57, 527-531.
Original article in IJSEM Online

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Pseudomonas diminuta Leifson and Hugh 1954, species.
Type strain: (see also StrainInfo.net) AJ 2067 = AS 1.1798 = ATCC 11568= CCEB 513 = BCRC (formerly CCRC) 11894 = CCUG 1187 = CIP 63.27 = DSM 7234 = IAM 12691 = IFO (now NBRC) 12697 = JCM 2788 = LMG 1793 = LMG 2089 = NCAIM B.01118 = NCCB 76050 = NCIMB 9393 = NCTC 8545 = VKM B-1297.
Sequence accession no. (16S rRNA gene) for the type strain: AB021415.
Etymology: L. adj. minutus, small; N.L. fem. adj. diminuta, defective, minute.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEIFSON (E.) and HUGH (R.): A new type of polar monotrichous flagellation. Journal of General Microbiology, 1954, 10, 68-70.

¤ Brevundimonas diminuta (Leifson and Hugh 1954) Segers et al. 1994, comb. nov.

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Pseudomonas donghuensis Gao et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) HYS=CCTCC AB 2012141=NRRL B-59108.
Sequence accession no. (whole genome shotgun sequencing project) for the type strain: AJJP00000000.
Whole-genome sequence accession no. for the type strain: AJJP00000000.
Etymology: N. L. fem. adj. dong. hu’ ensis, pertaining to Donghu, where the type strain was isolated.
Source: Environmental - freshwater.
Valid publication: Validation List no. 164. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 65 (2015), 2017-2025.
Validation List Online
Effective publication: Gao, J., Xie, G., Peng, F. and Xie, Z. 2015. Pseudomonas donghuensis sp. nov., exhibiting high-yields of siderophore. Antonie van Leeuwenhoek, 107, 83-94.
Effective publication Online

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Pseudomonas doudoroffii Baumann et al. 1972, species.
Type strain: (see also StrainInfo.net) ATCC 27123= CCUG 12373 = CCUG 16001 = CIP 74.9 = DSM 7028 = JCM 21046 = LMG 2180 = NBRC 103032.
Sequence accession no. (16S rRNA gene) for the type strain: AB019390.
Etymology: N.L. gen. masc. n. doudoroffii, of Doudoroff; named after M. Doudoroff.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: BAUMANN (L.), BAUMANN (P.), MANDEL (M.) and ALLEN (R.D.): Taxonomy of aerobic marine eubacteria. Journal of Bacteriology, 1972, 110, 402-429.

¤ Oceanimonas doudoroffii corrig. (Baumann et al. 1972) Brown et al. 2001, comb. nov.

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Pseudomonas duriflava Liu et al. 2008, sp. nov.
Type strain: (see also StrainInfo.net) HR2 = CGMCC 1.6858 = KCTC 22129.
Sequence accession no. (16S rRNA gene) for the type strain: EU046271.
Etymology: L. adj. durus, hard; L. adj. flavus, yellow; N.L. fem. adj. duriflava, hard yellow.
Valid publication: LIU (R.), LIU (H.), FENG (H.), WANG (X.), ZHANG (C.X.), ZHANG (K.Y.) and LAI (R.): Pseudomonas duriflava sp. nov., isolated from a desert soil. Int. J. Syst. Evol. Microbiol., 2008, 58, 1404-1408.
Original article in IJSEM Online

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Pseudomonas echinoides Heumann 1962, species.
Type strain: (see also StrainInfo.net) ATCC 14820= CCUG 2870 = CIP 103301 = DSM 50409 = DSM 1805 = ICPB 2835 = IFO (now NBRC) 15742 = LMG 7462 = NCIB (now NCIMB) 9420 = NRRL B-3126.
Sequence accession no. (16S rRNA gene) for the type strain: AB021370.
Etymology: Gr. n. echinos, hedgehog, sea-urchin; L. suff. -oides (from Gr. suff. -eides, from Gr. n. eidos, that which is seen, form, shape, figure), ressembling, similar; N.L. fem. adj. echinoides, prickly, like a hedgehog or a sea-urchin; intended to mean spiny shaped.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HEUMANN (W.): Die Methodik der Kreuzung sternbildender Bakterien. Biolgisches Zentralblatt, 1962, 81, 341-354.

¤ Sphingomonas echinoides (Heumann 1962) Denner et al. 1999, comb. nov.

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Pseudomonas elongata Humm 1946, species.
Type strain: (see also StrainInfo.net) ATCC 10144= DSM 6810 = LMG 2182 = NRRL B-762.
Sequence accession no. (16S rRNA gene) for the type strain: AF500006.
Etymology: L. fem. part. adj. elongata, elongated, stretched out.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HUMM (H.J.): Marine agar-digesting bacteria of the South Atlantic coast. Bulletin of Duke University Marine Station, 1946, 3, 45-75.

¤ Microbulbifer elongatus (Humm 1946) Yoon et al. 2003, comb. nov.

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Pseudomonas endophytica Ramírez-Bahena et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) BSTT44=LMG 28456=CECT 8691.
Sequence accession no. (16S rRNA) for the type strain: LN624760.
Etymology: Gr. pref. endo-, within; Gr. n. phyton, plant; L. fem. suff. -ica, adjectival suffix used with the sense of belonging to; N.L. fem. adj. endophytica, within the plant, endophytic.
Source: Plant.
Valid publication: RAMÍREZ-BAHENA, M. H., CUESTA, M. J., TEJEDOR, C., IGUAL, J. M., FERNÁNDEZ-PASCUAL, M. and PEIX, Á. 2015. Pseudomonas endophytica sp. nov., isolated from stem tissue of Solanum tuberosum L. in Spain. Int. J. Syst. Evol. Microbiol., 65, 2110-2117.
Original article in IJSEM Online

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Pseudomonas entomophila Mulet et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) L48 = CCUG 61470 = CECT 7985.
Sequence accession no. (16S rRNA gene) for the type strain: AY907566.
Etymology: Gr. n. entomon, insect; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. entomophila, insect loving, isolated from Drosophila melanogaster.
Valid publication: Validation List no. 146. Int. J. Syst. Evol. Microbiol., 2012, 62, 1443-1445.
Effective publication: MULET (M.), GOMILA (M.), LEMAITRE (B.), LALUCAT (J.) and GARCÍA-VALDÉS (E.): Taxonomic characterisation of Pseudomonas strain L48 and formal proposal of Pseudomonas entomophila sp. nov. Syst. Appl. Microbiol., 2012, 35, 145-149.
Validation List no. 146 in IJSEM Online - Effective publication Online

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Pseudomonas extremaustralis López et al. 2010, sp. nov.
Type strain: (see also StrainInfo.net) CT14-3 = CIP 109839 = DSM 17835.
Sequence accession no. (16S rRNA gene) for the type strain: AJ583501.
Etymology: L. adj. extremus, outermost, utmost, extreme; L. adj. australis, southern; N.L. fem. adj. extremaustralis, from the extreme South, the region of isolation (Far Southern Antarctica).
Valid publication: Validation List no. 132. Int. J. Syst. Evol. Microbiol., 2010, 60, 469-472.
Effective publication: LÓPEZ (N.I.), PETTINARI (M.J.), STACKEBRANDT (E.), TRIBELLI (P.M.), PÕTTER (M.), STEINBÜCHEL (A.) and MÉNDEZ (B.S.): Pseudomonas extremaustralis sp. nov., a poly(3-hydroxybutyrate) producer isolated from an Antarctic environment. Curr. Microbiol., 2009, 59, 514-519.
Validation List no. 132 in IJSEM Online - Effective publication Online

Note: In the abstract of the effective publication, the strain CT14-3 is cited as 14-3.
Reference: LISTS EDITOR, IJSEM: Validation List no. 132 (footnote ††). Int. J. Syst. Evol. Microbiol., 2010, 60, 469-472.
Validation List no. 132 in IJSEM Online

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Pseudomonas extremorientalis Ivanova et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) KMM 3447 = LMG 19695.
Sequence accession no. (16S rRNA gene) for the type strain: AF405328.
Etymology: L. adj. extremus, outermost, utmost, extreme; L. adj. orientalis, of or belonging to the East, Eastern; N.L. fem. adj. extremorientalis, of or belonging to the Far East of Russia, the region of isolation.
Valid publication: IVANOVA (E.P.), GORSHKOVA (N.M.), SAWABE (T.), HAYASHI (K.), KALINOVSKAYA (N.I.), LYSENKO (A.M.), ZHUKOVA (N.V.), NICOLAU (D.V.), KUZNETSOVA (T.A.), MIKHAILOV (V.V.) and CHRISTEN (R.): Pseudomonas extremorientalis sp. nov., isolated from a drinking water reservoir. Int. J. Syst. Evol. Microbiol., 2002, 52, 2113-2120.
Original article in IJSEM Online

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Pseudomonas facilis (Schatz and Bovell 1952) Davis 1969, species.
Type strain: (see also StrainInfo.net) ATCC 11228= CCUG 2113 = CFBP 2441 = CFBP 3608 = CIP 103302 = DSM 649 = LMG 2193 = NRRL B-4274.
Sequence accession no. (16S rRNA gene) for the type strain: AF078765.
Synonym: "Hydrogenomonas facilis" Schatz and Bovell 1952.
Etymology: L. fem. adj. facilis, easy, without difficulty.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SCHATZ (A.) and BOVELL (C.R.): Growth and hydrogenase activity of a new bacterium, Hydrogenomonas facilis. Journal of Bacteriology, 1952, 63, 87-98. 2 DAVIS (D.H.). In: DAVIS (D.H.), DOUDOROFF (M.), STANIER (R.Y.) and MANDEL (M.): Proposal to reject the genus Hydrogenomonas: taxonomic implications. International Journal of Systematic Bacteriology, 1969, 19, 375-390.

Note: On the Approved Lists of Bacterial Names and on the Approved Lists of Bacterial Names (Amended Edition), the authors of the publication in the International Journal of Systematic Bacteriology, are cited as DAVIS (D.H.), STANIER (R.Y.), DOUDOROFF (M.) and MANDEL (M.).

¤ Acidovorax facilis (Schatz and Bovell 1952) Willems et al. 1990, comb. nov.

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Pseudomonas ficuserectae Goto 1983, sp. nov.
Type strain: (see also StrainInfo.net) L7 = ATCC 35104= CCUG 32779 = CFBP 3224 = PDDCC 7848 = JCM 2400 = LMG 5694.
Sequence accession no. (16S rRNA gene) for the type strain: AB021378.
Etymology: L. gen. n. ficuserectae, of Ficus erecta, the name of the host species.
Valid publication: GOTO (M.): Pseudomonas ficuserectae sp. nov., the causal agent of bacterial leaf spot of Ficus erecta Thunb. Int. J. Syst. Bacteriol., 1983, 33, 546-550.
Original article in IJSEM Online

Note: Pseudomonas ficuserectae Goto 1983, ¤ Pseudomonas amygdali Psallidas and Panagopoulos 1975 (Approved Lists 1980), ¤ Pseudomonas meliae Ogimi 1981, and ¤ Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992 are members of a single genomospecies. Thus, these species should be considered synonymous and the correct name for this species should be Pseudomonas amygdali. Unfortunately, Pseudomonas amygdali, as delineated by DNA-DNA hybridization, could not be differentiated from other genomospecies by phenotypic tests. So the authors do not formally propose nomenclatural changes.
Reference: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Sutic and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

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Pseudomonas flava (Niklewski 1910) Davis 1969, species.
Type strain: (see also StrainInfo.net) ATCC 33667= CCUG 1658 = CCUG 22894 = CFBP 2438 = CIP 103271 = DSM 619 = JCM 21413 = LMG 2185 = NBRC 102514.
Sequence accession no. (16S rRNA gene) for the type strain: AF078771.
Synonym: "Hydrogenomonas flava" Niklewski 1910.
Etymology: L. fem. adj. flava, yellow.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 NIKLEWSKI (B.): Über die wasserstoffoxydation durch Mikroorganismen. Jahrbuch fur Wissenschaftliche Botanik, 1910, 48, 113-142. 2 DAVIS (D.H.). In: D.H. DAVIS, M. DOUDOROFF, R.Y. STANIER and M. MANDEL: Proposal to reject the genus Hydrogenomonas: taxonomic implications. International Journal of Systematic Bacteriology, 1969, 19, 375-390.

Note: On the Approved Lists of Bacterial Names and on the Approved Lists of Bacterial Names (Amended Edition), the authors of the publication in the International Journal of Systematic Bacteriology, are cited as DAVIS (D.H.), STANIER (R.Y.), DOUDOROFF (M.) and MANDEL (M.).

¤ Hydrogenophaga flava (Niklewski 1910) Willems et al. 1989, comb. nov.

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Pseudomonas flavescens Hildebrand et al. 1994, sp. nov.
Type strain: (see also StrainInfo.net) B62 = ATCC 51555= CCUG 49622 = CFBP 5586 = CIP 104204 = DSM 12071 = JCM 21586 = LMG 18387 = NBRC 103044 = NCPPB 3063.
Sequence accession no. (16S rRNA gene) for the type strain: U01916.
Etymology: L. part. adj. flavescens, becoming golden yellow.
Valid publication: HILDEBRAND (D.C.), PALLERONI (N.J.), HENDSON (M.), TOTH (J.) and JOHNSON (J.L.): Pseudomonas flavescens sp. nov., isolated from walnut blight cankers. Int. J. Syst. Bacteriol., 1994, 44, 410-415.
Original article in IJSEM Online

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Pseudomonas flectens Johnson 1956, species.
Type strain: (see also StrainInfo.net) ATCC 12775= CFBP 3281 = ICMP 745 = LMG 2187 = NCPPB 539.
Sequence accession no. (16S rRNA gene) for the type strain: AB021400.
Etymology: L. v. flectere, to bend, to curve; L. part. adj. flectens, bending.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: JOHNSON (J.C.): Pod twist: A previously unrecorded bacterial disease of French bean (Phaseolus vulgaris L.). Queensland Journal of Agricultural Science, 1956, 13, 127-158.

Note: Pseudomonas flectens Johnson 1956 (Approved Lists 1980) is not considered to be a member of the genus Pseudomonas sensu stricto.
References:
1 DE VOS (P.), GOOR (M.), GILLIS (M.) and DE LEY (J.): Ribosomal ribonucleic acid cistron similarities of phytopathogenic Pseudomonas species. Int. J. Syst. Bacteriol., 1985, 35, 169-184.
Original article in IJSEM Online
2 ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

¤ Phaseolibacter flectens (Johnson 1956) Halpern et al. 2013, comb. nov.

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Pseudomonas flexibilis Shin et al. 2016, comb. nov.
Type strain: (see also StrainInfo.net) ATCC 29606=LMG 29034.
Sequence accession no. (16S rRNA) for the type strain: GU269546.
Whole-genome sequence accession no. for the type strain: .
Basonym: ¤ Serpens flexibilis Hespell 1977 Approved Lists 1980.
Etymology: L. fem. adj. flexibilis, flexible, pliant.
Source: Environmental.
Valid publication: VALIDATION LIST no. 168. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 66 (2016), 1603-1606.
Validation List Online
Effective publication: Shin, S.-K., Hwang, C. Y., Cho, Y.-J. and Yi, H. 2015. Reclassification of Serpens flexibilis Hespell 1977 as Pseudomonas flexibilis comb. nov., with Pseudomonas tuomuerensis Xin et al. 2009 as a later heterotypic synonym. Syst. Appl. Microbiol. 38, 563-566.
Effective publication Online

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Pseudomonas fluorescens Migula 1895, species.
Type strain: (see also StrainInfo.net) RH 818 = M. Rhodes 28/5 = ATCC 13525= CCUG 1253 = CCEB 546 = CFBP 2102 = CIP 69.13 = DSM 50090 = IFO (now NBRC) 14160 = JCM 5963 = LMG 1794 = NCCB 76040 = NCIB (now NCIMB) 9046 = NCTC 10038 = NRRL B-14678 = VKM B-894.
Sequence accession no. (16S rRNA gene) for the type strain: D84013.
Synonyms: "Bacillus fluorescens liquefaciens" Flügge 1886, "Bacillus fluorescens" Trevisan 1889, "Bacterium fluorescens" (Trevisan 1889) Lehmann and Neumann 1896, "Liquidomonas fluorescens" (Trevisan 1889) Orla-Jensen 1909.
Etymology: N.L. part. adj. fluorescens, fluorescing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MIGULA (W.): Bacteriaceae (Stabchenbacterien). In: A. ENGLER and K. PRANTL (editors): Die Naturlichen Pfanzenfamilien, W. Engelmann, Leipzig, Teil I, Abteilung Ia, 1895, pp. 20-30.

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Pseudomonas formosensis Lin et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) CC-CY503=BCRC 80437=JCM 18415.
Sequence accession no. (16S rRNA) for the type strain: JF432053.
Etymology: N.L. fem. adj. formosensis, of or pertaining to Formosa (Taiwan), the beautiful island.
Source: Environmental.
Valid publication: LIN, S.-Y., HAMEED, A., LIU, Y.-C., HSU, Y.-H., LAI, W.-A. and YOUNG, C.-C. 2013. Pseudomonas formosensis sp. nov., a gamma-proteobacteria isolated from food-waste compost in Taiwan. Int. J. Syst. Evol. Microbiol., 63, 3168-3174.
Original article in IJSEM Online

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Pseudomonas fragi (Eichholz 1902) Gruber 1905, species.
Type strain: (see also StrainInfo.net) ATCC 4973= CCUG 556 = CFBP 4556 = CIP 55.4 = DSM 3456 = HAMBI 28 = IFO (now NBRC) 3458 = LMG 2191 = NCCB 69033 = NCTC 10689 = NRRL B-25 = NRRL B-727 = VKM B-898.
Sequence accession no. (16S rRNA gene) for the type strain: AF094733.
Synonym: "Bacterium fragi" Eichholz 1902.
Etymology: L. gen. n. fragi, of a strawberry-plant.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 EICHHOLZ (W.): Erdbeerbacillus (Bacterium fragi). Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung II, 1902, 9, 425-428. 2 GRUBER (T.): Ein weiterer Beiträg zür Aromabildung, speziell zür Bildung des Erdbeergeruches in der Gruppe "Pseudomonas", Pseudomonas fragariae II. Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung II, 1905, 14, 122-123.

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Pseudomonas frederiksbergensis Andersen et al. 2000, sp. nov.
Type strain: (see also StrainInfo.net) JAJ28 = ATCC BAA-257 = CIP 106887 = DSM 13022.
Sequence accession no. (16S rRNA gene) for the type strain: AJ249382.
Etymology: N.L. fem. adj. frederiksbergensis, of or belonging to to Frederiksberg near Copenhagen, Denmark, from where the organism was isolated.
Valid publication: ANDERSEN (S.M.), JOHNSEN (K.), SØRENSEN (J.), NIELSEN (P.) and JACOBSEN (C.S.): Pseudomonas frederiksbergensis sp. nov., isolated from soil at a coal gasification site. Int. J. Syst. Evol. Microbiol. 2000, 50, 1957-1964.
Original article in IJSEM Online

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Pseudomonas fulva Iizuka and Komagata 1963, species.
Type strain: (see also StrainInfo.net) ATCC 31418= CIP 106765 = CIP 106819 = IAM 1529 = IFO (now NBRC) 16637 = JCM 11242 = LMG 11722.
Sequence accession no. (16S rRNA gene) for the type strain: AB060136.
Etymology: L. fem. adj. fulva, tawny, deep yellow, yellowish brown.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IIZUKA (H.) and KOMAGATA (K.): New species of Pseudomonas belonged to fluorescent group (Studies on the microorganisms of cereal grains. Part V). Journal of the Agricultural Chemical Society of Japan, 1963, 37, 137-141.

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Pseudomonas fuscovaginae (ex Tanii et al. 1976) Miyajima et al. 1983, sp. nov., nom. rev.
Type strain: (see also StrainInfo.net) 6801 = CCUG 32780 = CFBP 2065 = CIP 106695 = DSM 7231 = ICMP 5940 = LMG 2158 = NCPPB 3085 = PDDCC 5940.
Sequence accession no. (16S rRNA gene) for the type strain: FJ483519.
Synonym: "Pseudomonas fuscovaginae" Tanii et al. 1976.
Etymology: L. adj. fuscus, tawny, fuscous; L. n. vagina, vagina, sheath; N.L. gen. n. fuscovaginae, of a fuscous vagina.
Valid publication: MIYAJIMA (K.), TANII (A.) and AKITA (T.): Pseudomonas fuscovaginae sp. nov., nom. rev. Int. J. Syst. Bacteriol., 1983, 33, 656-657.
Original article in IJSEM Online

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Pseudomonas gelidicola Kadota 1951, species.
Type strain: (see also StrainInfo.net) CIP 106748 = IAM 1127 = JCM 20065 = LMG 21612.
Sequence accession no. (16S rRNA gene) for the type strain: not found.
Etymology: L. adj. gelidus, ice-cold, very cold, icy; L. suff. -cola (from L. n. incola), inhabitant, dweller; N.L. n. gelidicola, ice-dweller.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: KADOTA (H.): Studies on the biochemical activities of marine bacteria. I. On the agar-decomposing bacteria in the sea. Memoirs of the College of Science, Kyoto University, 1951, 59, 54-67.

Note: The type strain is no longer available in CIP, DSMZ, JCM (IAM), and LMG.
References: YARZA (P.): Personal communication (March 04, 2012).

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Pseudomonas geniculata (Wright 1895) Chester 1901, species.
Type strain: (see also StrainInfo.net) ATCC 19374= JCM 13324 = LMG 2195 = NCIB (now NCIMB) 9428.
Sequence accession no. (16S rRNA gene) for the type strain: AB021404.
Synonym: "Bacillus geniculatus" Wright 1895.
Etymology: L. fem. adj. geniculata, jointed.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 WRIGHT (J.H.): Report on the results of an examination of the water supply of Philadelphia. Memoirs of the National Academy of Science, 1895, 7, 422-484. 2 CHESTER (F.D.): A Manual of Determinative Bacteriology. The Macmillan Co., New York, 1901, pp. 1-401.

Note: According to Anzai et al., Pseudomonas geniculata (Wright 1895) Chester 1901 (Approved Lists 1980) does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

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Pseudomonas gessardii Verhille et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) CIP 105469 = CFML 95-251.
Sequence accession no. (16S rRNA gene) for the type strain: AF074384.
Etymology: N.L. gen. masc. n. gessardii, of Gessard, named after C. Gessard, French chemist who isolated "Bacterium aeruginosum" for the first time in 1882 and studied its pigment.
Valid publication: VERHILLE (S.), BAÏDA (N.), DABBOUSSI (F.), HAMZE (M.), IZARD (D.) and LECLERC (H.): Pseudomonas gessardii sp. nov. and Pseudomonas migulae sp. nov., two new species isolated from natural mineral waters. Int. J. Syst. Bacteriol., 1999, 49, 1559-1572.
Original article in IJSEM Online

Pseudomonas gessardii Verhille et al. 1999 emend. van den Beld et al. 2016.
Reference: VAN DEN BELD, M. J., REINDERS, E., NOTERMANS, D. W. and REUBSAET, F. A. 2016. Possible misidentification of species in the Pseudomonas fluorescens lineage as Burkholderia pseudomallei and Francisella tularensis, and emended descriptions of Pseudomonas brenneri, Pseudomonas gessardii and Pseudomonas proteolytica. Int. J. Syst. Evol. Microbiol., 66, 3420-3425.
Original article.

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Pseudomonas gladioli Severini 1913, species.
Type strain: (see also StrainInfo.net) ATCC 10248= CCUG 1782 = CFBP 2427 = CIP 105410 = DSM 4285 = HAMBI 2157 = ICMP 3950 = IFO (now NBRC) 13700 = JCM 9311 = LMG 2216 = NCCB 38018 = NCPPB 1891 = NCTC 12378 = NRRL B-793.
Sequence accession no. (16S rRNA gene) for the type strain: EU024168.
Etymology: L. n. gladiolus, a small sword, lily, and also a botanical generic name; L. gen. n. gladioli, of lily, of Gladiolus.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: SEVERINI (G.): Una bacteriosi dell' Ixia maculata e del Gladiolus colvilli. Annali di Botanica (Roma), 1913, 11, 413-424.

¤ Burkholderia gladioli (Severini 1913) Yabuuchi et al. 1993, comb. nov.

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Pseudomonas glareae Romanenko et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) KMM 9500=NRIC 0939.
Sequence accession no. (16S rRNA) for the type strain: LC011944.
Whole-genome sequence accession no. for the type strain: -.
Etymology: L. gen. n. glareae, of pebble stones.
Source: Environmental - marine.
Valid publication: Validation List no. 166. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 65 (2015), 3763-3767.
Validation List Online
Effective publication: Romanenko, L.A., Tanaka, N., Svetashev, V.I. and Mikhailov, V.V. 2015. Pseudomonas glareae sp. nov., a marine sediment-derived bacterium with antagonistic activity. Arch. Microbiol. 197, 693-699.
Effective publication Online
Note: Etymology corrected on validation..

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Pseudomonas glathei Zolg and Ottow 1975, species.
Type strain: (see also StrainInfo.net) ATCC 29195= CFBP 4791 = CIP 105421 = DSM 50014 = JCM 10563 = LMG 14190.
Sequence accession no. (16S rRNA gene) for the type strain: U96935.
Etymology: N.L. gen. masc. n. glathei, of Glathe, named after H. Glathe of Giessen, Germany.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: ZOLG (W.) and OTTOW (J.C.G.): Pseudomonas glathei sp. nov., a new nitrogen-scavenging rod isolated from acid lateritic relicts in Germany. Zeitschrift fur Allgemeine Mikrobiologie, 1975, 15, 287-299.

¤ Burkholderia glathei (Zolg and Ottow 1975) Vandamme et al. 1997, comb. nov.

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Pseudomonas glumae Kurita and Tabei 1967, species.
Type strain: (see also StrainInfo.net) ATCC 33617= CCUG 20835 = CFBP 4900 = CIP 106418 = DSM 9512 = ICMP 3655 = LMG 2196 = NCPPB 2981 = NIAES 1169.
Sequence accession no. (16S rRNA gene) for the type strain: U96931.
Etymology: L. n. gluma, a hull or husk; L. gen. n. glumae, of a husk.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: KURITA (T.) and TABEI (H.): On the pathogenic bacterium of bacterial grain rot of rice. Annals of the Phytopathological Society, Japan, 1967, 33, 111.

¤ Burkholderia glumae (Kurita and Tabei 1967) Urakami et al. 1994, comb. nov.

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Pseudomonas graminis Behrendt et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) P 294/08 = CCUG 51504 = CIP 105897 = DSM 11363.
Sequence accession no. (16S rRNA gene) for the type strain: Y11150.
Etymology: L. n. gramen, grass; L. gen. n. graminis, of grass, the source of the organism.
Valid publication: BEHRENDT (U.), ULRICH (A.), SCHUMANN (P.), ERLER (W.), BURGHARDT (J.) and SEYFARTH (W.): A taxonomic study of bacteria isolated from grasses: a proposed new species Pseudomonas graminis sp. nov. Int. J. Syst. Bacteriol., 1999, 49, 297-308.
Original article in IJSEM Online

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Pseudomonas granadensis Pascual et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) F-278,770=DSM 28040=LMG 27940.
Sequence accession no. (16S rRNA) for the type strain: HG764746.
Etymology: N.L. fem. adj. granadensis, denizen of the province of Granada in south-east Spain, where the type strain was isolated.
Source: Environmental - soil.
Valid publication: PASCUAL, J., GARCÍA-LÓPEZ, M., BILLS, G. F. and GENILLOUD, O. 2015. Pseudomonas granadensis sp. nov., a new bacterial species isolated from the Tejeda, Almijara and Alhama Natural Park, Granada, Spain. Int. J. Syst. Evol. Microbiol., 65, 625-632
Original article in IJSEM Online

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Pseudomonas grimontii Baïda et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) ATCC BAA-140 = CFML 97-514 = CIP 106645.
Sequence accession no. (16S rRNA gene) for the type strain: AF268029.
Etymology: N.L. gen. masc. n. grimontii, of Grimont, in honour of the French bacteriologist P. A. D. Grimont, for all his work in the field of bacterial taxonomy.
Valid publication: BAÏDA (N.), YAZOURH (A.), SINGER (E.) and IZARD (D.): Pseudomonas grimontii sp. nov. Int. J. Syst. Evol. Microbiol., 2002, 52, 1497-1503.
Original article in IJSEM Online

Note: Pseudomonas grimontii Baïda et al. 2002 was previously known as "fluorescent Pseudomonas group 97-514".

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Pseudomonas guangdongensis Yang et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) SgZ-6=CCTCC AB 2012022=KACC 16606.
Sequence accession no. (16S rRNA) for the type strain: JX274436.
Etymology: N.L. fem. adj. guangdongensis, of or pertaining to Guangdong, a province in south-east China, from where the type strain was isolated.
Source: Industrial.
Valid publication: YANG, G., HAN, L., WEN, J. and ZHOU, S. 2013. Pseudomonas guangdongensis sp. nov., isolated from an electroactive biofilm, and emended description of the genus Pseudomonas Migula 1894. Int. J. Syst. Evol. Microbiol., 63, 4599-4605.
Original article in IJSEM Online

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Pseudomonas guariconensis Toro et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) PCAVU11=LMG 27394=CECT 8262.
Sequence accession no. (16S rRNA) for the type strain: HF674459.
Etymology: N.L. fem. adj. guariconensis, of or belonging to Guárico, the Venezuelan state where the type strain was isolated.
Source: Environmental - soil.
Reference: TORO, M., RAMÍREZ-BAHENA, M.-H., CUESTA, M. J., VELÁZQUEZ, E. and PEIX, A. 2013. Pseudomonas guariconensis sp. nov., isolated from rhizospheric soil. Int. J. Syst. Evol. Microbiol., 63, 4413-4420.
Original article in IJSEM Online

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Pseudomonas guineae Bozal et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) M8 = CECT 7231 = LMG 24016.
Sequence accession no. (16S rRNA gene) for the type strain: AM491810.
Etymology: N.L. gen. masc. n. guineae, of Guinea, in honour of the late Professor Jesús Guinea, a prominent Spanish microbiologist, who isolated this strain.
Valid publication: BOZAL (N.), MONTES (M.J.) and MERCADÉ (E.): Pseudomonas guineae sp. nov., a novel psychrotolerant bacterium from an Antarctic environment. Int. J. Syst. Evol. Microbiol., 2007, 57, 2609-2612.
Original article in IJSEM Online

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Pseudomonas halophila Fendrich 1989, sp. nov.
Type strain: (see also StrainInfo.net) II = DSM 3050.
Sequence accession no. (16S rRNA gene) for the type strain: AB021383.
Etymology: Gr. n. hals halos, salt; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. halophila, salt-loving.
Valid publication: Validation List no. 29. Int. J. Syst. Bacteriol., 1989, 39, 205-206.
Effective publication: FENDRICH (C.): Halovibrio variabilis gen. nov. sp. nov., Pseudomonas halophila sp. nov. and a new halophilic aerobic coccoid Eubacterium from Great Salt Lake, Utah, USA. Syst. Appl. Microbiol., 1988, 11, 36-43.
Validation List no. 29 in IJSEM Online - Effective publication Online

Note: According to Anzai et al., Pseudomonas halophila Fendrich 1989 does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

Note:
During the course of studies on halophilic, Gram-negative bacteria, a comparison of two strains of halophilic bacteria, Pseudomonas halophila DSM 3050 (type strain of the species) and ¤ Halomonas variabilis (basonym ¤ Halovibrio variabilis) DSM 3051 (type strain of the species) demonstrated that the characteristics of strain DSM 3050 corresponded to the original description of ¤ Halomonas variabilis (basonym ¤ Halovibrio variabilis) and those of DSM 3051 to Pseudomonas halophila [1].
Based on data collected, Sorokin and Tindall 2006 [2] conclude that DSM 3051 is clearly a member of the genus ¤ Halomonas, and that DSM 3050 clearly is not a member of the genus ¤ Halomonas.
In a Request for an Opinion, Sorokin and Tindall 2006 [2] propose that the following steps should be undertaken by the Judicial Comission.
(i) The type of the name ¤ Halovibrio variabilis Fendrich 1989 is to be regarded as DSM 3050, not as currently published DSM 3051.
(ii) The circumscription of the genus Halovibrio should be emended as proposed by Sorokin et al. 2006.
(iii) This would also have the effect of placing members of the genus Halovibrio outside of the genus ¤ Halomonas and would mean that the names ¤ Halovibrio variabilis and ¤ Halomonas variabilis cannot be treated as synonyms.
(iv) DSM 3051 should be recognized as the type strain of Pseudomonas halophila.
(v) The species name ¤ Halomonas variabilis (Fendrich 1989) Dobson and Franzmann 1996 is to be rejected, because it was created in the belief that ¤ Halovibrio variabilis was correctly represented by DSM 3051.
(vi) Recognition of DSM 3051 as the type strain of Pseudomonas halophila would mean that data collected for this strain indicate that it should be properly placed (at least at present) within the genus ¤ Halomonas. However, the name Halomonas halophila already exists [¤ Halomonas halophila (Quesada et al. 1984) Dobson and Franzmann 1996] and it would be necessary to create a new name (nomen novum) to reflect this taxonomic interpretation. Sorokin and Tindall propose the combination "Halomonas utahensis" nom. nov. (homotypic synonym Pseudomonas halophila) to accommodate strain DSM 3051, with the current authors (and date) being the authority of the name. The type strain is DSM 3051.
(vii) Given the special circumstances under which the data were collected and published, Sorokin and Tindall 2006 also request that the Judicial Commission recognize that the species name Halovibrio denitrificans was validly published by Sorokin et al. 2006.
References:
1 SOROKIN (D.Y.), TOUROVA (T.P.), GALINSKI (E.A.), BELLOCH (C.) and TINDALL (B.J.): Extremely halophilic denitrifying bacteria from hypersaline inland lakes, Halovibrio denitrificans sp. nov. and Halospina denitrificans gen. nov., sp. nov., and evidence that the genus name Halovibrio Fendrich 1989 with the type species Halovibrio variabilis should be associated with DSM 3050. Int. J. Syst. Evol. Microbiol., 2006, 56, 379-388.
Original article by Sorokin et al. 2006 in IJSEM Online
2 SOROKIN (D.Y.) and TINDALL (B.J.): The status of the genus name Halovibrio Fendrich 1989 and the identity of the strains Pseudomonas halophila DSM 3050 and Halomonas variabilis DSM 3051. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2006, 56, 487-489.
Original article by Sorokin and Tindall 2006 in IJSEM Online

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Pseudomonas helleri Von Neubeck et al. 2016, sp. nov.
Type strain: (see also StrainInfo.net) WS4917=DSM 29165=LMG 28433.
Sequence accession no. (16S rRNA) for the type strain: KP738715.
Whole-genome sequence accession no. for the type strain: -.
Etymology: N.L. gen. masc. n. helleri, named in honour of Knut Heller, former microbiologist of the Max-Rubner-Institute in Kiel, Germany, with great merit in food microbiology.
Source: Animal.
Valid publication: VON NEUBECK, M., HUPTAS, C., GLUCK, C., KREWINKEL, M., STOECKEL, M., STRESSLER, T., FISCHER, L., HINRICHS, J., SCHERER, S. and WENNING, M. 2016. Pseudomonas helleri sp. nov. and Pseudomonas weihenstephanensis sp. nov., isolated from raw cow's milk. Int. J. Syst. Evol. Microbiol., 66, 1163-1173.
Original article in IJSEM Online

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Pseudomonas helmanticensis Ramírez et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) OHA11=LMG 28168=CECT 8548.
Sequence accession no. (16S rRNA) for the type strain: HG940537.
Etymology: N.L. fem. adj. helmanticensis, pertaining to Helmantica, the name of Salamanca in Roman times.
Source: Environmental - soil.
Valid publication: RAMÍREZ-BAHENA, M.-H., CUESTA, M. J., FLORES-FÉLIX, J. D., MULAS, R., RIVAS, R. L., CASTRO-PINTO, J., BRAÑAS, J., MULAS, D., GONZÁLEZ-ANDRÉS, F., VELÁZQUEZ, E. and PEIX, Á. 2014. Pseudomonas helmanticensis sp. nov., isolated from forest soil. Int. J. Syst. Evol. Microbiol., 64, 2338-2345.
Original article in IJSEM Online

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Pseudomonas hibiscicola Moniz 1963, species.
Type strain: (see also StrainInfo.net) ATCC 19867= ICMP 3945 = JCM 13361 = LMG 980 = NCPPB 1683.
Sequence accession no. (16S rRNA gene) for the type strain: AB021405.
Etymology: N.L. n. Hibiscus, a botanical generic name; L. suff. -cola (from L. n. incola), inhabitant, dweller; N.L. n. hibiscicola, Hibiscus-dweller (pathogenic to Hibiscus Rosa-sinensis).
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MONIZ (L.): Leaf spot of apple blossom. Current Science, 1963, 32, 177.

Note: Pseudomonas hibiscicola Moniz 1963 (Approved Lists 1980) and ¤ Pseudomonas beteli corrig. (Ragunathan 1928) Savulescu 1947 (Approved Lists 1980) are misnamed and should be transferred to the genus ¤ Xanthomonas Dowson 1939 (Approved Lists 1980). Pseudomonas hibiscicola, Pseudomonas beteli corrig. and ¤ Xanthomonas maltophilia (Hugh 1981) Swings et al. 1983 are heterotypic synonyms. According to the Bacteriological Code (1990 revision) Pseudomonas hibiscicola and Xanthomonas maltophilia should be renamed Xanthomonas beteli because beteli is the oldest specific epithet. The authors do not feel however the need to formally propose nomenclatural changes.
Reference: VAN DEN MOOTER (M.) and SWINGS (J.): Numerical analysis of 295 phenotypic features of 266 Xanthomonas strain and related strains and an improved taxonomy of the genus. Int. J. Syst. Bacteriol., 1990, 40, 348-369.
Original article in IJSEM Online

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Pseudomonas hussainii Hameed et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) CC-AMH-11=JCM 19513=BCRC 80696.
Sequence accession no. (16S rRNA) for the type strain: KF582606.
Etymology: N.L. masc. gen. n. hussainii, named after Syed Abdulla Hussain (13 August 1944-30 December 2009), an Indian ornithologist and avian gut biologist.
Source: Animal.
Valid publication: HAMEED, A., SHAHINA, M., LIN, S.-Y., LIU, Y.-C. and YOUNG, C.-C. 2014. Pseudomonas hussainii sp. nov., isolated from droppings of a seashore bird, and emended descriptions of Pseudomonas pohangensis, Pseudomonas benzenivorans and Pseudomonas segetis. Int. J. Syst. Evol. Microbiol., 64, 2330-2337.
Original article in IJSEM Online

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Pseudomonas huttiensis Leifson 1962, species.
Type strain: (see also StrainInfo.net) ATCC 14670= CIP 103296 = DSM 10281 = IAM 14941 = JCM 21423 = LMG 2199.
Sequence accession no. (16S rRNA gene) for the type strain: AB021366.
Etymology: N.L. fem. adj. huttiensis, pertaining to Lower Hutt, New Zealand.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEIFSON (E.): The bacterial flora of distilled and stored water. III. New species of the genera Corynebacterium, Flavobacterium, Spirillum and Pseudomonas. International Bulletin of Bacteriological Nomenclature and Taxonomy, 1962, 12, 161-170.

¤ Herbaspirillum huttiense (Leifson 1962) Ding and Yokota 2004, comb. nov.

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Pseudomonas indica Pandey et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) IMT37 = DSM 14015 = JCM 21544 = MTCC 3713 = NBRC 103045.
Sequence accession no. (16S rRNA gene) for the type strain: AF302795.
Etymology: L. fem. adj. indica, pertaining to India, where the organism was isolated.
Valid publication: PANDEY (K.K.), MAYILRAJ (S.) and CHAKRABARTI (T.): Pseudomonas indica sp. nov., a novel butane-utilizing species. Int. J. Syst. Evol. Microbiol., 2002, 52, 1559-1567.
Original article in IJSEM Online

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Pseudomonas indigofera (Voges 1893) Migula 1900, species.
Type strain: (see also StrainInfo.net) ATCC 19706= CCUG 2873 = CCUG 32860 = CIP 103306 = DSM 3303 = LMG 6867.
Sequence accession no. (16S rRNA gene) for the type strain: AB021385.
Synonym: "Bacillus indigoferus" Voges 1893.
Etymology: N.L. n. indigo (from Fr. n. indigo, derived from L. n. indicum, indigo) the dye indigo; L. suff. -fer -fera -ferum (from L. v. fero, to bear), bearing; N.L. fem. adj. indigofera, bearing indigo.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 VOGES (O.): Über einige im Wasser vorkommende Pigmentbakterien. Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung I, 1893, 14, 301-314. 2 MIGULA (W.): System der Bakterien, Vol. 2, Gustav Fischer, Jena, 1900, pp. 1-1068.

¤ Vogesella indigofera (Voges 1893) Grimes et al. 1997, comb. nov.

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Pseudomonas iners Iizuka and Komagata 1964, species.
Type strain: (see also StrainInfo.net) CIP 106746 = IAM 1419 = NBRC 102196.
Sequence accession no. (16S rRNA gene) for the type strain: AB021408.
Etymology: L. fem. adj. iners, inactive, inert.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IIZUKA (H.) and KOMAGATA (K.): Microbiological studies on petroleum and natural gas. II. Determination of pseudomonads isolated from oil-brines and related materials. Journal of General and Applied Microbiology, 1964, 10, 223-231.

¤ Marinobacterium georgiense González et al. 1997 emend. Satomi et al. 2002.

Note: Pseudomonas iners and ¤ Marinobacterium georgiense are unified as ¤ Marinobacterium georgiense González et al. 1997 emend. Satomi et al. 2002 [Rule 44].

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Pseudomonas japonica Pungrasmi et al. 2008, sp. nov.
Type strain: (see also StrainInfo.net) WL = IAM 15071 = JCM 21532 = TISTR 1526.
Sequence accession no. (16S rRNA gene) for the type strain: AB126621.
Etymology: N.L. fem. adj. japonica, pertaining to Japan, from where the isolate originated.
Valid publication: Validation List no. 122. Int. J. Syst. Evol. Microbiol., 2008, 58, 1511-1512.
Effective publication: PUNGRASMI (W.), LEE (H.S.), YOKOTA (A.) and OHTA (A.): Pseudomonas japonica sp. nov., a novel species that assimilates straight chain alkylphenols. J. Gen. Appl. Microbiol., 2008, 54, 61-69.
Validation List no. 122 in IJSEM Online - Effective publication Online

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Pseudomonas jessenii Verhille et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) ATCC 700870= CCUG 42059 = CFML 95-307 = CIP 105274.
Sequence accession no. (16S rRNA gene) for the type strain: AF068259.
Etymology: N.L. masc. gen. n. jessenii, of Jessen, named after O. Jessen, eminent Danish bacteriologist who contributed substantially to the knowledge of fluorescent pseudomonads.
Valid publication: Validation List no. 70. Int. J. Syst. Bacteriol., 1999, 49, 935-936.
Effective publication: VERHILLE (S.), BAIDA (N.), DABBOUSSI (F.), IZARD (D.) and LECLERC (H.): Taxonomic study of bacteria isolated from natural mineral waters: proposal of Pseudomonas jessenii sp. nov. and Pseudomonas mandelii sp. nov. Syst. Appl. Microbiol., 1999, 22, 45-58.
Validation List no. 70 in IJSEM Online - Effective publication Online

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Pseudomonas jinjuensis Kwon et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) Pss 26 = JCM 21621 = KACC 10760 = LMG 21316 = NBRC 103047.
Sequence accession no. (16S rRNA gene) for the type strain: AF468448.
Etymology: N.L. adj. jinjuensis, referring to Jinju Region in Korea, where the bacteria were first found.
Valid publication: KWON (S.W.), KIM (J.S.), PARK (I.C.), YOON (S.H.), PARK (D.H.), LIM (C.K.) and GO (S.J.): Pseudomonas koreensis sp. nov., Pseudomonas umsongensis sp. nov. and Pseudomonas jinjuensis sp. nov., novel species from farm soils in Korea. Int. J. Syst. Evol. Microbiol., 2003, 53, 21-27.
Original article in IJSEM Online

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Pseudomonas kilonensis Sikorski et al. 2001, sp. nov.
Type strain: (see also StrainInfo.net) 520-20 = CFBP 5372 = DSM 13647 = JCM 11939.
Sequence accession no. (16S rRNA gene) for the type strain: AJ292426.
Etymology: L. fem. adj. kilonensis (after Civium Kilonensis, the ancient Latin name of the town of Kiel, Germany), of or belonging to Kiel.
Valid publication: SIKORSKI (J.), STACKEBRANDT (E.) and WACKERNAGEL (W.): Pseudomonas kilonensis sp. nov., a bacterium isolated from agricultural soil. Int. J. Syst. Evol. Microbiol., 2001, 51, 1549-1555.
Original article in IJSEM Online

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Pseudomonas knackmussii Stolz et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) B13 = DSM 6978 = LMG 23759.
Sequence accession no. (16S rRNA gene) for the type strain: AF039489.
Etymology: N.L. masc. gen. n. knackmussii, of Knackmuss, in honour of Hans-Joachim Knackmuss, who initiated the biochemical studies about the degradation of chlorinated aromatics and other xenobiotic compounds by strain B13T and various other micro-organisms.
Valid publication: STOLZ (A.), BUSSE (H.J.) and KÄMPFER (P.): Pseudomonas knackmussii sp. nov. Int. J. Syst. Evol. Microbiol., 2007, 57, 572-576.
Original article in IJSEM Online

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Pseudomonas koreensis Kwon et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) Ps 9-14 = JCM 14769 = KACC 10848 = LMG 21318.
Sequence accession no. (16S rRNA gene) for the type strain: AF468452.
Etymology: N.L. fem. adj. koreensis, pertaining to Korea.
Valid publication: KWON (S.W.), KIM (J.S.), PARK (I.C.), YOON (S.H.), PARK (D.H.), LIM (C.K.) and GO (S.J.): Pseudomonas koreensis sp. nov., Pseudomonas umsongensis sp. nov. and Pseudomonas jinjuensis sp. nov., novel species from farm soils in Korea. Int. J. Syst. Evol. Microbiol., 2003, 53, 21-27.
Original article in IJSEM Online

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Pseudomonas kunmingensis Xie et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) HL22-2=CGMCC 1.12273=DSM 25974.
Sequence accession no. (16S rRNA) for the type strain: JQ246444.
Whole-genome sequence accession no. for the type strain: N/A.
Etymology: N.L. fem. adj. kunmingensis, pertaining to Kunming, the region in Yunnan, China, where the type strain was isolated.
Source: Environmental.
Valid publication: XIE, F., MA, H., QUAN, S., LIU, D., CHEN, G., CHAO, Y. and QIAN, S. 2014. Pseudomonas kunmingensis sp. nov., an exopolysaccharide-producing bacterium isolated from a phosphate mine. Int. J. Syst. Evol. Microbiol., 64, 559-564.
Original article in IJSEM Online

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Pseudomonas kuykendallii Hunter and Manter 2012, sp. nov.
Type strain: (see also StrainInfo.net) H2 = LMG 26364 = NRRL B-59562.
Sequence accession no. (16S rRNA gene) for the type strain: JF749828.
Etymology: N.L. gen. masc. n. kuykendallii, of Kuykendall, in honor of L. David Kuykendall for his contributions to the field of microbiology.
Valid publication: Validation List no. 148. Int. J. Syst. Evol. Microbiol., 2012, 62, 2549-2554.
Effective publication: HUNTER (W.J.) and MANTER (D.K.): Pseudomonas kuykendallii sp. nov.: a novel γ-proteobacteria isolated from a hexazinone degrading bioreactor. Curr. Microbiol., 2012, 65, 170-175.
Validation List no. 148 in IJSEM Online - Effective publication Online

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Pseudomonas lanceolata Leifson 1962, species.
Type strain: (see also StrainInfo.net) ATCC 14669= CCUG 48175 = CFBP 5587 = CIP 103297 = IAM 14947 = NBRC 103051.
Sequence accession no. (16S rRNA gene) for the type strain: AB021390.
Etymology: L. fem. adj. lanceolata, lanceolated, lancet-shaped.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEIFSON (E.): The bacterial flora of distilled and stored water. III. New species of the genera Corynebacterium, Flavobacterium, Spirillum and Pseudomonas. International Bulletin of Bacteriological Nomenclature and Taxonomy, 1962, 12, 161-170.

¤ Curvibacter lanceolatus (Leifson 1962) Ding and Yokota 2004, comb. nov.

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Pseudomonas lemoignei Delafield et al. 1965, species.
Type strain: (see also StrainInfo.net) ATCC 17989= CCUG 2114 = CIP 103794 = DSM 7445 = LMG 2207 = NCTC 10937.
Sequence accession no. (16S rRNA gene) for the type strain: X92555.
Etymology: N.L. gen. masc. n. lemoignei, of Lemoigne; named after M.H. Lemoigne, a French bacteriologist.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DELAFIELD (F.P.), DOUDOROFF (M.), PALLERONI (N.J.), LUSTY (C.J.) and CONTOPOULOU (C.R.): Decomposition of poly-β-hydroxybutyrate by pseudomonads. Journal of Bacteriology, 1965, 90, 1455-1466.

¤ Paucimonas lemoignei (Delafield et al. 1965) Jendrossek 2001, comb. nov.

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Pseudomonas libanensis Dabboussi et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) CCUG 43190 = CFML 96-195 = CIP 105460.
Sequence accession no. (16S rRNA gene) for the type strain: AF057645.
Etymology: L. n. Libanus, Lebanon; N.L. fem. adj. libanensis, pertaining to Lebanon.
Valid publication: DABBOUSSI (F.), HAMZE (M.), ELOMARI (M.), VERHILLE (S.), BAIDA (N.), IZARD (D.) and LECLERC (H.): Pseudomonas libanensis, sp. nov., a new species isolated from Lebanese spring waters. Int. J. Syst. Bacteriol, 1999, 49, 1091-1101.
Original article in IJSEM Online

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Pseudomonas lini Delorme et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) DLE411J = CCUG 51522 = CFBP 5737 = ICMP 14138.
Sequence accession no. (16S rRNA gene) for the type strain: AY035996.
Etymology: L. n. linum, flax; L. gen. n. lini, of flax, referring to the plant genus, Linum, from which the type strain CFBP 5737T was isolated.
Valid publication: DELORME (S.), LEMANCEAU (P.), CHRISTEN (R.), CORBERAND (T.), MEYER (J.M.) and GARDAN (L.): Pseudomonas lini sp. nov., a novel species from bulk and rhizospheric soils. Int. J. Syst. Evol. Microbiol., 2002, 52, 513-523.
Original article in IJSEM Online

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Pseudomonas linyingensis He et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) LYBRD3-7=CGMCC 1.10701=LMG 25967.
Sequence accession no. (16S rRNA) for the type strain: HM246142.
Whole-genome sequence accession no. for the type strain: -.
Etymology: -.
Source: Environmental - soil.
Valid publication: Validation List no. 163. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 65 (2015), 1105-1111.
Validation List Online
Effective publication: HE, W.-H., WANG, Y.-N., DU, X., ZHOU, Y., JIA, B., BIAN, J., LIU, S.-J. and CHEN, G.-C. 2012. Pseudomonas linyingensis sp. nov.: a novel bacterium isolated from wheat soil subjected to long-term herbicides application. Curr Microbiol, 65, 595-600.
Effective publication Online

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Pseudomonas litoralis Pascual et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) 2SM5 = CECT 7670 = KCTC 23093.
Sequence accession no. (16S rRNA gene) for the type strain: FN908483.
Etymology: L. fem. adj. litoralis, of or belonging to the seashore.
Valid publication: PASCUAL (J.), LUCENA (T.), RUVIRA (M.A.), GIORDANO (A.), GAMBACORTA (A.), GARAY (E.), ARAHAL (D.R.), PUJALTE (M.J.) and MACIÁN (M.C.): Pseudomonas litoralis sp. nov., isolated from Mediterranean seawater. Int. J. Syst. Evol. Microbiol., 2012, 62, 438-444.
Original article in IJSEM Online

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Pseudomonas lundensis Molin et al. 1986, sp. nov.
Type strain: (see also StrainInfo.net) 138 of Molin and Ternström = 573 = ATCC 49968= CCM 3503 = CCUG 18757 = CFBP 4564 = CIP 103272 = DSM 6252 = LMG 13517 = VKM B-2183.
Sequence accession no. (16S rRNA gene) for the type strain: AB021395.
Etymology: N.L. fem. adj. lundensis, pertaining to the city of Lund,
Reference: MOLIN (G.), TERNSTRÖM (A.) and URSING (J.): Pseudomonas lundensis, a new bacterial species isolated from meat. Int. J. Syst. Bacteriol., 1986, 36, 339-342.
Original article in IJSEM Online

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Pseudomonas lurida Behrendt et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) P 513/18 = DSM 15835 = LMG 21995.
Sequence accession no. (16S rRNA gene) for the type strain: AJ581999.
Etymology: L. fem. adj. lurida, yellowish, to indicate the yellow-greenish fluorescent pigment of the organism.
Valid publication: BEHRENDT (U.), ULRICH (A.), SCHUMANN (P.), MEYER (J.M.) and SPRÖER (C.): Pseudomonas lurida sp. nov., a fluorescent species associated with the phyllosphere of grasses. Int. J. Syst. Evol. Microbiol., 2007, 57, 979-985.
Original article in IJSEM Online

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Pseudomonas lutea Peix et al. 2004, sp. nov.
Type strain: (see also StrainInfo.net) OK2 = CECT 5822 = LMG 21974.
Sequence accession no. (16S rRNA gene) for the type strain: AY364537.
Etymology: L. fem. adj. lutea, yellow, referring to the yellowish pigment produced by this bacterium.
Valid publication: PEIX (A.), RIVAS (R.), SANTA-REGINA (I.), MATEOS (P.F.), MARTÍNEZ-MOLINA (E.), RODRÍGUEZ-BARRUECO (C.) and VELÁZQUEZ (E.): Pseudomonas lutea sp. nov., a novel phosphate-solubilizing bacterium isolated from the rhizosphere of grasses. Int. J. Syst. Evol. Microbiol., 2004, 54, 847-850.
Original article in IJSEM Online

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Pseudomonas luteola Kodama et al. 1985, sp. nov.
Type strain: (see also StrainInfo.net) KS0921 = G.L. Gilardi 4239 = ATCC 43273= CCUG 37974 = CFBP 3007 = CIP 102995 = DSM 6975 = IAM 13000 = JCM 3352 = LMG 7041 = NBRC 103146.
Sequence accession no. (16S rRNA gene) for the type strain: D84002.
Etymology: L. fem. dim. adj. luteola, yellowish.
Valid publication: KODAMA (K.), KIMURA (N.) and KOMAGATA (K.): Two new species of Pseudomonas: P. oryzihabitans isolated from rice paddy and clinical specimens and P. luteola isolated from clinical specimens. Int. J. Syst. Bacteriol., 1985, 35, 467-474.
Original article in IJSEM Online

Note: According to Holmes et al. 1987, Pseudomonas luteola Kodama et al. 1985 is an earlier heterotypic synonym of ¤ Chryseomonas polytricha Holmes et al. 1986.
Reference: HOLMES (B.) STEIGERWALT (A.G.), WEAVER (R.E.) and BRENNER (D.J.): Chryseomonas luteola comb. nov. and Flavimonas oryzihabitans gen. nov., comb. nov., Pseudomonas-like species from human clinical specimens and formerly known, respectively, as groups Ve-1 and Ve-2. Int. J. Syst. Bacteriol., 1987, 37, 245-250.
Original article in IJSEM Online

¤ Chryseomonas luteola (Kodama et al. 1985) Holmes et al. 1987, comb. nov.

Note: Pseudomonas luteola Kodama et al. 1985 is an earlier heterotypic synonym of ¤ Chryseomonas polytricha Holmes et al. 1986. The type strain of Pseudomonas luteola was 78% related to the type strain of Chryseomonas polytricha Holmes et al. 1986. Holmes et al. 1987 propose the new combinaison Chryseomonas luteola (Kodama et al. 1985) Holmes et al. 1987 for this organism.
Reference: HOLMES (B.) STEIGERWALT (A.G.), WEAVER (R.E.) and BRENNER (D.J.): Chryseomonas luteola comb. nov. and Flavimonas oryzihabitans gen. nov., comb. nov., Pseudomonas-like species from human clinical specimens and formerly known, respectively, as groups Ve-1 and Ve-2. Int. J. Syst. Bacteriol., 1987, 37, 245-250.
Original article in IJSEM Online

Pseudomonas luteola Kodama et al. 1985.
Type strain: (see also StrainInfo.net) KS0921 = G.L. Gilardi 4239 = ATCC 43273= CCUG 37974 = CIP 102995 = DSM 6975 = IAM 13000 = JCM 3352.
Sequence accession no. (16S rRNA gene) for the type strain: D84002.
Etymology:
Reference: ANZAI (Y.), KUDO (Y.) and OYAIZU (H.): The phylogeny of the genera Chryseomonas, Flavimonas, and Pseudomonas supports synonymy of these three genera. Int. J. Syst. Bacteriol., 1997, 47, 249-251.
Original article in IJSEM Online

Note: The level of similarity between the 16S rRNA sequence of the strain KS0921 = G.L. Gilardi 4239 = IAM 13000 = JCM 3352 (the type strain of the species Chryseomonas luteola (Kodama et al. 1985) Holmes et al. 1987) and the 16S rRNA of the strain DSM 50071 = ATCC 10145(the type strain of the species Pseudomonas aeruginosa (Schroeter 1872) Migula 1900 (Approved Lists 1980) is 94.0%. This result suggests that Chryseomonas Holmes et al. 1986 emend. Holmes et al. 1987 is a later synonym of Pseudomonas Migula 1894 (Approved Lists 1980) and that the name Pseudomonas luteola should be used for this species.

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Pseudomonas mallei (Zopf 1885) Redfearn et al. 1966, species.
Type strain: (see also StrainInfo.net) ATCC 23344= NCTC 12938.
Sequence accession no. (16S rRNA gene) for the type strain: AF110188.
Synonyms: "Bacillus mallei" Zopf 1885, "Pfeifferella mallei" (Zopf 1885) Buchanan 1918, "Malleomyces mallei" (Zopf 1885) Pribram 1933, "Actinobacillus mallei" (Zopf 1885) Thompson 1933, "Loefferella mallei" (Zopf 1885) Holden 1935, "Acinetobacter mallei" (Zopf 1885) Steel and Cowan 1964.
Etymology: L. n. malleus, a hammer, maul, the disease of glanders; L. gen. n. mallei, of glanders.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 ZOPF (W.): Die Spaltpilze, 3rd ed. Edward Trewendt, Breslau, 1885, pp. 1-127. 2 REDFEARN (M.S.), PALLERONI (N.J.) and STANIER (R.Y.): A comparative study of Pseudomonas pseudomallei and Bacillus mallei. Journal of General Microbiology, 1966, 43, 293-313.

¤ Burkholderia mallei (Zopf 1885) Yabuuchi et al. 1993, comb. nov.

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Pseudomonas maltophilia (ex Hugh and Ryschenkow 1961) Hugh 1981, sp. nov., nom. rev.
Type strain: (see also StrainInfo.net) Stanier 67 = Hugh 810-2 = RH 1168 = AS 1.1788 = ATCC 13637= CCUG 5866 = CFBP 3035 = CCM 1640 = BCRC (formerly CCRC) 10737 = CIP 60.77 = DSM 50170 = IAM 12423 = ICMP 17033 = IFO (now NBRC) 14161 = IMET 10402 = JCM 1975 = LMG 958 = NCAIM B.01119 = NCCB 68018 = NCIMB 9203 = NCPPB 1974 = NCTC 10257 = NRC 729 = NRRL B-2756 = VKM B-591.
Sequence accession no. (16S rRNA gene) for the type strain: AB294553.
Synonym: "Pseudomonas maltophilia" Hugh and Ryschenkow 1961.
Etymology: N.L. n. maltum, malt; Gr. n. philia, friendship; N.L. n. maltophilia, intended to mean friend of malt.
Valid publication: HUGH (R.): Pseudomonas maltophilia sp. nov., nom. rev. Int. J. Syst. Bacteriol., 1981, 31, 195.
Original article in IJSEM Online

Note: The name Pseudomonas maltophilia is also documented as Hugh and Ryschenkow 1960 [1]. However, the effective publication is a paper by Hugh and Ryschenkow 1961 [2].
References:
1 HUGH (R.) and RYSCHENKOW (E.): An alcaligenes-like Pseudomonas species. Bacteriol. Proc. 1960, p. 78.
2 HUGH (R.) and RYSCHENKOW (E.): Pseudomonas maltophilia, an alcaligenes-like species. J. Gen. Microbiol., 1961, 26, 123-132.
Original article Online

¤ Xanthomonas maltophilia (Hugh 1981) Swings et al. 1983 emend. van den Mooter and Swings 1990.

¤ Stenotrophomonas maltophilia (Hugh 1981) Palleroni and Bradbury 1993, comb. nov.

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Pseudomonas mandelii Verhille et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) ATCC 700871= CCUG 42058 = CFML 95-303 = CIP 105273 = JCM 21619 = NBRC 103147.
Sequence accession no. (16S rRNA gene) for the type strain: AF058286.
Etymology: N.L. masc. gen. n. mandelii, of Mandel, dedicated to M. Mandel, American bacteriologist.
Valid publication: Validation List no. 70. Int. J. Syst. Bacteriol., 1999, 49, 935-936.
Effective publication: VERHILLE (S.), BAIDA (N.), DABBOUSSI (F.), IZARD (D.) and LECLERC (H.): Taxonomic study of bacteria isolated from natural mineral waters: proposal of Pseudomonas jessenii sp. nov. and Pseudomonas mandelii sp. nov. Syst. Appl. Microbiol., 1999, 22, 45-58.
Validation List no. 70 in IJSEM Online - Effective publication Online

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Pseudomonas marginalis (Brown 1918) Stevens 1925, species.
Type strain: (see also StrainInfo.net) ATCC 10844= CFBP 1387 = CFBP 2037 = CIP 106712 = DSM 13124 = ICMP 3553 = LMG 2215 = NCPPB 667.
Sequence accession no. (16S rRNA gene) for the type strain: Z76663.
Synonym: "Bacterium marginale" Brown 1918.
Etymology: L. n. margo marginis, edge, margin; L. fem. suff. -alis, suffix denoting pertaining to; N.L. fem. adj. marginalis, marginal.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 BROWN (N.A.): Some bacterial diseases of lettuce. Journal of Agricultural Research, 1918, 13, 367-388. 2 STEVENS (F.L.) (ed.): Plant Disease Fungi, MacMillan Co., New York, 1925, pp. 469.

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Pseudomonas marina (Cobet et al. 1970) Baumann et al. 1972, species.
Type strain: (see also StrainInfo.net) 219 = ATCC 25374= CCUG 49558 = CCUG 49558 = CIP 104765 = DSM 4741 = LMG 2217 = NBRC 102605.
Sequence accession no. (16S rRNA gene) for the type strain: AJ306890.
Synonym: "Arthrobacer marinus" Cobet et al. 1970.
Etymology: L. fem. adj. marina, of or belonging to the sea, marine.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 COBET (A.B.), WIRSEN (C.) and JONES (G.E.): The effect of nickel on a marine bacterium, Arthrobacter marinus sp. nov. Journal of General Microbiology, 1970, 62, 159-169. 2 BAUMANN (L.), BAUMANN (P.), MANDEL (M.) and ALLEN (R.D.): Taxonomy of aerobic marine eubacteria. Journal of Bacteriology, 1972, 110, 402-429.

¤ Deleya marina (Cobet et al. 1970) Baumann et al. 1983, comb. nov.

¤ Halomonas marina (Cobet et al. 1970) Dobson and Franzmann 1996, comb. nov.

¤ Cobetia marina (Cobet et al. 1970) Arahal et al. 2002, comb. nov.

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Pseudomonas marincola Romanenko et al. 2008, sp. nov.
Type strain: (see also StrainInfo.net) JCM 14761 = KMM 3042 = NRIC 0729.
Sequence accession no. (16S rRNA gene) for the type strain: AB301071.
Etymology: L. n. mare, the sea; L. n. incola, inhabitant; N.L. n. marincola, inhabitant of the sea.
Valid publication: ROMANENKO (L.A.), UCHINO (M.), TEBO (B.M.), TANAKA (N.), FROLOVA (G.M.) and MIKHAILOV (V.V.): Pseudomonas marincola sp. nov., isolated from marine environments. Int. J. Syst. Evol. Microbiol., 2008, 58, 706-710.
Original article in IJSEM Online

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Pseudomonas matsuisoli Lin et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) CC-MHH0089=BCRC 80771=JCM 30078.
Sequence accession no. (16S rRNA) for the type strain: KJ720680.
Etymology: L. neut. n. solum, soil; N.L. gen. n. matsuisoli, of soil of Matsu Island, Taiwan, where the type strain was isolated.
Source: Environmental - soil.
Valid publication: LIN, S.-Y., HAMEED, A., HUNG, M.-H., LIU, Y.-C., HSU, Y.-H., YOUNG, L.-S. and YOUNG, C.-C. 2015. Pseudomonas matsuisoli sp. nov., isolated from a soil sample. Int. J. Syst. Evol. Microbiol., 65, 902-909.
Original article in IJSEM Online

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Pseudomonas mediterranea Catara et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFBP 5447 = ICMP 14184.
Sequence accession no. (16S rRNA gene) for the type strain: AF386080.
Etymology: L. fem. adj. mediterranea, mediterranean, referring to the fact that most strains of this species were isolated in Mediterranean countries.
Valid publication: CATARA (V.), SUTRA (L.), MORINEAU (A.), ACHOUAK (W.), CHRISTEN (R.) and GARDAN (L.): Phenotypic and genomic evidence for the revision of Pseudomonas corrugata and proposal of Pseudomonas mediterranea sp. nov. Int. J. Syst. Evol. Microbiol., 2002, 52, 1749-1758.
Original article in IJSEM Online

Note: Pseudomonas mediterranea Catara et al. 2002 was previously known as ¤ Pseudomonas corrugata phenon B.

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Pseudomonas meliae Ogimi 1981, sp. nov.
Type strain: (see also StrainInfo.net) No. 2 = ATCC 33050= CFBP 3225 = ICMP 6289 = LMG 2220 = NCPPB 3033.
Sequence accession no. (16S rRNA gene) for the type strain: not found.
Etymology: N.L. n. Melia (from Gr. n. melia, the ash), a botanical genus name; N.L. gen. n. meliae, of Melia, isolated from Melia azedarach L. (Chinaberry tree).
Valid publication: Validation List no. 7. Int. J. Syst. Bacteriol., 1981, 31, 382-383.
Effective publication: OGIMI (C.): Studies on bacterial gall of chinaberry (Melia Azedarach Lin.), caused by Pseudomonas meliae n. sp. Bull Coll. Agr., Univ of the Ryukyus, 1977, 24, 497-556.
Validation List no. 7 in IJSEM Online

Note: Pseudomonas meliae Ogimi 1981, ¤ Pseudomonas amygdali Psallidas and Panagopoulos 1975 (Approved Lists 1980), ¤ Pseudomonas ficuserectae Goto 1983, and ¤ Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992 are members of a single genomospecies. Thus, these species should be considered synonymous and the correct name for this species should be Pseudomonas amygdali. Unfortunately, Pseudomonas amygdali, as delineated by DNA-DNA hybridization, could not be differentiated from other genomospecies by phenotypic tests. So the authors do not formally propose nomenclatural changes.
Reference: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Sutic and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

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Pseudomonas mendocina Palleroni 1970, species.
Type strain: (see also StrainInfo.net) ATCC 25411= CCUG 1781 = CFBP 2434 = CIP 75.21 = DSM 50017 = IFO (now NBRC) 14162 = JCM 5966 = LMG 1223 = NCCB 76043 = NCTC 10897 = VKM B-972.
Sequence accession no. (16S rRNA gene) for the type strain: D84016.
Etymology: N.L. fem. adj. mendocina, pertaining to Mendoza (Argentina).
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: PALLERONI (N.J.). In: PALLERONI (N.J.), DOUDOROFF (M.), STANIER (R.Y.), SOLANES (R.E.) and MANDEL (M.): Taxonomy of the aerobic pseudomonads, the properties of the Pseudomonas stutzeri group. Journal of General Microbiology, 1970, 60, 215-231.

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Pseudomonas mephitica Claydon and Hammer 1939, species.
Type strain: (see also StrainInfo.net) ATCC 33665= CCUG 2513 = CIP 106720 = JCM 13308 = NCIB (now NCIMB) 9672.
Sequence accession no. (16S rRNA gene) for the type strain: AM748811.
Etymology: L. fem. adj. mephitica, mephitic, pestilential.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: CLAYDON (T.J.) and HAMMER (B.W.): A skunk-like odor of bacterial origin in butter. Journal of Bacteriology, 1939, 37, 251-258.

Notes:
1 In the Approved Lists of Bacterial Names and in the Approved Lists of Bacterial Names Amended Edition, the pages number of the paper by Claydon and Hammer is erroneously cited as 252-258.
2 According to Anzai et al., Pseudomonas mephitica Claydon and Hammer 1939 (Approved Lists 1980) does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

¤ Janthinobacterium lividum (Eisenberg 1891) De Ley et al. 1978 (Approved Lists 1980).

Note: According to Kämpfer et al. 2008, Pseudomonas mephitica Claydon and Hammer 1939 (Approved Lists 1980) is a later heterotypic synonym of ¤ Janthinobacterium lividum (Eisenberg 1891) De Ley et al. 1978 (Approved Lists 1980).
Reference: KÄMPFER (P.), FALSEN (E.) and BUSSE (H.J.): Reclassification of Pseudomonas mephitica Claydon and Hammer 1939 as a later heterotypic synonym of Janthinobacterium lividum (Eisenberg 1891) De Ley et al. 1978. Int. J. Syst. Evol. Microbiol., 2008, 58, 136-138.
Original article in IJSEM Online

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Pseudomonas meridiana Reddy et al. 2004, sp. nov.
Type strain: (see also StrainInfo.net) CMS 38 = DSM 15319 = MTCC 4993.
Sequence accession no. (16S rRNA gene) for the type strain: AJ537602.
Etymology: L. fem. adj. meridiana, of or belonging to the south or south side, southern, southerly, meridional; pertaining to the South Pole.
Valid publication: REDDY (G.S.N.), MATSUMOTO (G.I.), SCHUMANN (P.), STACKEBRANDT (E.) and SHIVAJI (S.): Psychrophilic pseudomonads from Antarctica: Pseudomonas antarctica sp. nov., Pseudomonas meridiana sp. nov. and Pseudomonas proteolytica sp. nov. Int. J. Syst. Evol. Microbiol., 2004, 54, 713-719.
Original article in IJSEM Online

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Pseudomonas mesophilica Austin and Goodfellow 1979, species.
Type strain: (see also StrainInfo.net) A47 = ATCC 29983= CCUG 16482 = CIP 101129 = DSM 1708 = ICPB 4095 = IFO (now NBRC) 15688 = LMG 5275 = NCIB (now NCIMB) 11561 = JCM 2829.
Sequence accession no. (16S rRNA gene) for the type strain: AB175636.
Etymology: Gr. adj. mesos, middle; Gr. adj. philikos -ê -on, friendly; N.L. fem. adj. mesophilica, friendly to the middle, because of its preference for moderate temperatures.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: AUSTIN (B.) and GOODFELLOW (M.): Pseudomonas mesophilica, a new species of pink bacteria isolated from leaf surfaces. International Journal of Systematic Bacteriology, 1979, 29, 373-378.

¤ Methylobacterium mesophilicum (Austin and Goodfellow 1979) Green and Bousfield 1983, comb. nov.

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Pseudomonas migulae Verhille et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) CCUG 43165 = CIP 105470 = CFML 95-321 = JCM 21628 = NBRC 103157.
Sequence accession no. (16S rRNA gene) for the type strain: AF074383.
Etymology: N.L. masc. gen. n. migulae, of Migula, named after W. Migula, who created the generic name Pseudomonas.
Valid publication: VERHILLE (S.), BAÏDA (N.), DABBOUSSI (F.), HAMZE (M.), IZARD (D.) and LECLERC (H.): Pseudomonas gessardii sp. nov. and Pseudomonas migulae sp. nov., two new species isolated from natural mineral waters. Int. J. Syst. Bacteriol., 1999, 49, 1559-1572.
Original article in IJSEM Online

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Pseudomonas mixta Bowman et al. 1989, sp. nov.
Type strain: (see also StrainInfo.net) ACM 1762 = ATCC 49108= CCUG 35206 = CIP 104068 = DSM 4832 = LMG 11547 = UQM 1762.
Sequence accession no. (16S rRNA gene) for the type strain: X65589.
Etymology: L. fem. part. adj. mixta (from L. v. misceo), mixed, referring to mixed flagellation.
Valid publication: Validation List no. 29. Int. J. Syst. Bacteriol., 1989, 39, 205-206.
Effective publication: BOWMAN (J.P.), SLY (L.I.) and HAYWARD (A.C.): Pseudomonas mixta sp. nov., a bacterium from soil with degradative activity on a variety of complex polysaccharides. Syst. Appl. Microbiol., 1988, 11, 53-69.
Validation List no. 29 in IJSEM Online - Effective publication Online

¤ Telluria mixta (Bowman et al. 1989) Bowman et al. 1993, comb. nov.

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Pseudomonas mohnii Cámara et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) IpA-2 = CCUG 53115 = DSM 18327.
Sequence accession no. (16S rRNA gene) for the type strain: AM293567.
Etymology: N.L. gen. masc. n. mohnii, of Mohn, named in honour of William Mohn, for his pioneering contributions to the study of the biodegradation of resin acids.
Valid publication: CÁMARA (B.), STRÖMPL (C.), VERBARG (S.), SPRÖER (C.), PIEPER (D.H.) and TINDALL (B.J.): Pseudomonas reinekei sp. nov., Pseudomonas moorei sp. nov. and Pseudomonas mohnii sp. nov., novel species capable of degrading chlorosalicylates or isopimaric acid. Int. J. Syst. Evol. Microbiol., 2007, 57, 923-931.
Original article in IJSEM Online

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Pseudomonas monteilii Elomari et al. 1997, sp. nov.
Type strain: (see also StrainInfo.net) ATCC 700476= CCUG 38736 = CFML 90-60 = CIP 104883 = DSM 14164 = JCM 21585 = NBRC 103158.
Sequence accession no. (16S rRNA gene) for the type strain: AF064458.
Etymology: N.L. masc. gen. n. monteilii, of Monteil, in honor of Henri Monteil, a French microbiologist.
Valid publication: ELOMARI (M.), COROLER (L.), VERHILLE (S.), IZARD (D.) and LECLERC (H.): Pseudomonas monteilii sp. nov., isolated from clinical specimens. Int. J. Syst. Bacteriol., 1997, 47, 846-852.
Original article in IJSEM Online

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Pseudomonas moorei Cámara et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) RW10 = CCUG 53114 = DSM 12647.
Sequence accession no. (16S rRNA gene) for the type strain: AM293566.
Etymology: N.L. gen. masc. n. moorei, of Moore, named in honour of Edward R. B. Moore, for his important contributions to the study of the genus Pseudomonas.
Valid publication: CÁMARA (B.), STRÖMPL (C.), VERBARG (S.), SPRÖER (C.), PIEPER (D.H.) and TINDALL (B.J.): Pseudomonas reinekei sp. nov., Pseudomonas moorei sp. nov. and Pseudomonas mohnii sp. nov., novel species capable of degrading chlorosalicylates or isopimaric acid. Int. J. Syst. Evol. Microbiol., 2007, 57, 923-931.
Original article in IJSEM Online

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Pseudomonas moraviensis Tvrzová et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) 1B4 = CCM 7280 = DSM 16007 = JCM 14770.
Sequence accession no. (16S rRNA gene) for the type strain: AY970952.
Etymology: N.L. fem. adj. moraviensis, pertaining to Moravia, the region of the Czech Republic where the type strain 1B4 was isolated.
Valid publication: TVRZOVÁ (L.), SCHUMANN (P.), SPRÖER (C.), SEDLÁČEK (I.), PÁČOVÁ (Z.), ŠEDO (O.), ZDRÁHAL (Z.), STEFFEN (M.) and LANG (E.): Pseudomonas moraviensis sp. nov. and Pseudomonas vranovensis sp. nov., soil bacteria isolated on nitroaromatic compounds, and emended description of Pseudomonas asplenii. Int. J. Syst. Evol. Microbiol., 2006, 56, 2657-2663.
Original article in IJSEM Online

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Pseudomonas mosselii Dabboussi et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFML 90-83 = ATCC BAA-99 = CIP 105259.
Sequence accession no. (16S rRNA gene) for the type strain: AF072688.
Etymology: N.L. masc. gen. n. mosselii, of Mossel, in honour of David A. A. Mossel, a Dutch microbiologist, for his contribution to medical and food microbiology.
Valid publication: DABBOUSSI (F.), HAMZE (M.), SINGER (E.), GEOFFROY (V.), MEYER (J.M.) and IZARD (D.): Pseudomonas mosselii sp. nov., a novel species isolated from clinical specimens. Int. J. Syst. Evol. Microbiol., 2002, 52, 363-376.
Original article in IJSEM Online

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Pseudomonas mucidolens Levine and Anderson 1932, species.
Type strain: (see also StrainInfo.net) ATCC 4685= CCUG 1424 = CIP 103298 = JCM 2781 = LMG 2223 = NBRC 103159 = NCTC 8068 = NRRL B-16.
Sequence accession no. (16S rRNA gene) for the type strain: D84017.
Etymology: L. adj. mucidus, musty; L. v. oleo, to emit a smell, to smell of; N.L. part. adj. mucidolens, musty smelling.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEVINE (M.) and ANDERSON (D.Q.): Two new species of bacteria causing mustiness in eggs. Journal of Bacteriology, 1932, 23, 337-347.

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Pseudomonas multiresinivorans Mohn et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) IpA-1 = ATCC 700690= CCUG 51655 = CIP 106706 = NBRC 103160.
Sequence accession no. (16S rRNA gene) for the type strain: X96787.
Etymology: L. adj. multus, many; L. fem. n. resina, resin; L. part. adj. vorans, eating, devouring; N.L. part. adj. multiresinivorans, devouring many resins.
Valid publication: Validation List no. 70. Int. J. Syst. Bacteriol., 1999, 49, 935-936.
Effective publication: MOHN (W.W.), WILSON (A.E.), BICHO (P.) and MOORE (E.R.B.): Physiological and phylogenetic diversity of bacteria growing on resin acids. Syst. Appl. Microbiol., 1999, 22, 68-78.
Validation List no. 70 in IJSEM Online - Effective publication Online

¤ Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980) emend. Lang et al. 2007.

Note: According to Lang et al. 2007 Pseudomonas multiresinivorans Mohn et al. 1999 is a later heterotypic synonym of ¤ Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980) emend. Lang et al. 2007.
Reference: LANG (E.), GRIESE (B.), SPRÖER (C.), SCHUMANN (P.), STEFFEN (M.) and VERBARG (S.): Characterization of 'Pseudomonas azelaica' DSM 9128, leading to emended descriptions of Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) and Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980), including Pseudomonas multiresinivorans as its later heterotypic synonym. Int. J. Syst. Evol. Microbiol., 2007, 57, 878-882.
Original article in IJSEM Online

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Pseudomonas nautica Baumann et al. 1972, species.
Type strain: (see also StrainInfo.net) ATCC 27132= CCUG 16007 = DSM 50418 = LMG 2226 = NBRC 103161.
Sequence accession no. (16S rRNA gene) for the type strain: EU373087.
Etymology: L. fem. adj. nautica, nautical.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: BAUMANN (L.), BAUMANN (P.), MANDEL (M.) and ALLEN (R.D.): Taxonomy of aerobic marine eubacteria. Journal of Bacteriology, 1972, 110, 402-429.

¤ Marinobacter hydrocarbonoclasticus Gauthier et al. 1992.

Note: A polyphasic taxonomic approach revealed that the type strains of Pseudomonas nautica Bauman et al. 1972 (Approved Lists 1980) and ¤ Marinobacter hydrocarbonoclasticus Gauthier et al. 1992 were highly similar and should be considered members of the same species. The authors propose the unification of the two species Pseudomonas nautica and Marinobacter hydrocarbonoclasticus as Marinobacter hydrocarbonoclasticus.
References: SPRÖER (C.), LANG (E.), HOBECK (P.), BURGHARDT (J.), STACKEBRANDT (E.) and TINDALL (B.J.): Transfer of Pseudomonas nautica to Marinobacter hydrocarbonoclasticus. Int. J. Syst. Bacteriol., 1998, 48, 1445-1448.
Original article in IJSEM Online

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Pseudomonas nitritireducens Wang et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) WZBFD3-5A2=CGMCC 1.10702=LMG 25966.
Sequence accession no. (16S rRNA) for the type strain: HM246143.
Whole-genome sequence accession no. for the type strain: -.
Etymology: -.
Source: Environmental - soil.
Valid publication: Validation List no. 163. List of new names and new combinations previously effectively, but not validly, published. Int. J. Syst. Evol. Microbiol., 65 (2015), 1105-1111.
Validation List Online
Effective publication: WANG, Y.-N., HE, W.-H., HE, H., DU, X., JIA, B., ZENG, Z.-P., AN, M.-L. and CHEN, G.-C. 2012. Pseudomonas nitritireducens sp. nov., a nitrite reduction bacterium isolated from wheat soil. Arch Microbiol, 194, 809-813.
Effective publication Online

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Pseudomonas nitroreducens Iizuka and Komagata 1964, species.
Type strain: (see also StrainInfo.net) P-6 = ATCC 33634= CCUG 12538 = CIP 106747 = DSM 14399 = IAM 1439 = IFO (now NBRC) 12694 = JCM 2782.
Sequence accession no. (16S rRNA gene) for the type strain: AM088473.
Etymology: Gr. n. nitron, nitre, nitrate; L. part. adj. reducens, drawing backwards, bringing back to a state or condition; N.L. part. adj. nitroreducens, nitrate reducing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IIZUKA (H.) and KOMAGATA (K.): Microbiological studies on petroleum and natural gas. I. Determination of hydrocarbon-utilizing bacteria. Journal of General and Applied Microbiology, 1964, 10, 207-221.

Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980) emend. Lang et al. 2007.
Type strain: (see also StrainInfo.net) P-6 = ATCC 33634= CCUG 12538 = CIP 106747 = DSM 14399 = IAM 1439 = JCM 2782 = NBRC 12694.
Sequence accession no. (16S rRNA gene) for the type strain: AM088473.
Etymology: Gr. n. nitron, nitre, nitrate; L. part. adj. reducens, drawing backwards, bringing back to a state or condition; N.L. part. adj. nitroreducens, nitrate reducing.
Reference: LANG (E.), GRIESE (B.), SPRÖER (C.), SCHUMANN (P.), STEFFEN (M.) and VERBARG (S.): Characterization of 'Pseudomonas azelaica' DSM 9128, leading to emended descriptions of Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) and Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980), including Pseudomonas multiresinivorans as its later heterotypic synonym. Int. J. Syst. Evol. Microbiol., 2007, 57, 878-882.
Original article in IJSEM Online

Notes:
1 According to Lang et al. 2007 Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas multiresinivorans Mohn et al. 1999.
Reference: LANG (E.), GRIESE (B.), SPRÖER (C.), SCHUMANN (P.), STEFFEN (M.) and VERBARG (S.): Characterization of 'Pseudomonas azelaica' DSM 9128, leading to emended descriptions of Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) and Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980), including Pseudomonas multiresinivorans as its later heterotypic synonym. Int. J. Syst. Evol. Microbiol., 2007, 57, 878-882.
Original article in IJSEM Online
2 Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980) emend. Lang et al. 2007 encompasses the strain DSM 9128, previously known as "Pseudomonas azelaica" Janota-Bassalik et al., 1971.
References:
1 LANG (E.), GRIESE (B.), SPRÖER (C.), SCHUMANN (P.), STEFFEN (M.) and VERBARG (S.): Characterization of 'Pseudomonas azelaica' DSM 9128, leading to emended descriptions of Pseudomonas citronellolis Seubert 1960 (Approved Lists 1980) and Pseudomonas nitroreducens Iizuka and Komagata 1964 (Approved Lists 1980), including Pseudomonas multiresinivorans as its later heterotypic synonym. Int. J. Syst. Evol. Microbiol., 2007, 57, 878-882.
Original article in IJSEM Online
2 Janota-Bassalik (L.), Bohdanowicz-Strucinska (B.) and Noras (A.): Observations on Pseudomonas sp. highly tolerant to creosote, isolated from railway wood sleepers. Acta. Microbiol. Pol. B, 1971, 3, 143-153.

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Pseudomonas oceani Wang and Sun 2016, sp. nov.
Type strain: (see also StrainInfo.net) KX 20=CGMCC 1.15195=DSM 100277.
Sequence accession no. (16S rRNA) for the type strain: KT449732.
Etymology: L. gen. n. oceani, of the ocean.
Source: Environmental - marine.
Valid publication: WANG, M. Q. and SUN, L. 2016. Pseudomonas oceani sp. nov., isolated from deep seawater. Int. J. Syst. Evol. Microbiol., 66, 4250-4255.
Original article in IJSEM Online

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Pseudomonas oleovorans Lee and Chandler 1941, species.
Type strain: (see also StrainInfo.net) ATCC 8062= CCUG 2087 = CFBP 5589 = CIP 59.11 = DSM 1045 = IFO (now NBRC) 13583 = JCM 11598 (IAM 1508) = LMG 2229 = NCIMB 6576 = NCTC 10692 = NRRL B-778 = VKM B-1522.
Sequence accession no. (16S rRNA gene) for the type strain: D84018.
Etymology: L. n. oleum, oil; L. part. adj. vorans, eating, devouring; N.L. part. adj. oleovorans, oil devouring.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEE (M.) and CHANDLER (A.C.): A study of the nature, growth and control of bacteria in cutting compounds. Journal of Bacteriology, 1941, 41, 373-386.

Notes:
1 Pseudomonas oleovorans is isolated from oil-water emulsions used as lubricants and cooling agents in the cutting and grinding of metals. Apparently the organism lives on some normal constituent of the cutting compound, probably the naphthenic acids, which act as emulsifying agents. Consequently, the name oleovorans may not be appropriate.
Reference: PALLERONI (N.J.). In: D.J. BRENNER, N.R. KRIEG, J.T. STALEY and G. M. GARRITY (editors), Bergey's Manual of Systematic Bacteriology, second edition, vol. 2 (The Proteobacteria), part B (The Gammaproteobacteria), Springer, New York, 2005, pp. 323-379.
2 According to Saha et al. 2010, Pseudomonas oleovorans Lee and Chandler 1941 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas pseudoalcaligenes Stanier 1966 (Approved Lists 1980).
References: List of Changes in Taxonomic Opinion no. 12. Int. J. Syst. Evol. Microbiol., 2010, 60, 1482-1483 [SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.]
List of Changes in Taxonomic Opinion no. 12 in IJSEM Online - Effective publication Online

Pseudomonas oleovorans Lee and Chandler 1941 emend. Saha et al. 2010.
Type strain: (see also StrainInfo.net) ATCC 8062= CCUG 2087 = CFBP 5589 = CIP 59.11 = DSM 1045 = IFO (now NBRC) 13583 = JCM 11598 (IAM 1508) = LMG 2229 = NCIMB 6576 = NCTC 10692 = NRRL B-778 = VKM B-1522.
Sequence accession no. (16S rRNA gene) for the type strain: D84018.
Etymology: L. n. oleum, oil; L. part. adj. vorans, eating, devouring; N.L. part. adj. oleovorans, oil devouring.
References: List of Changes in Taxonomic Opinion no. 12. Int. J. Syst. Evol. Microbiol., 2010, 60, 1482-1483 [SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.]
List of Changes in Taxonomic Opinion no. 12 in IJSEM Online - Effective publication Online

Note: Subsequently, this species has been divided into subspecies (see: below).

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Pseudomonas oleovorans subsp. lubricantis Saha et al. 2010, subsp. nov.
Type strain: (see also StrainInfo.net) RS1 = ATCC BAA-1494 = DSM 21016.
Sequence accession no. (16S rRNA gene) for the type strain: DQ842018.
Etymology: L. v. lubricare, to lubricate; N.L. n lubricans -antis (from L. part. adj. lubricans) a lubricant; N.L. gen. n. lubricantis, of/from lubricating oil, the source of the organism.
Valid publication: Validation List no. 134. Int. J. Syst. Evol. Microbiol., 2010, 60, 1477-1479.
Effective publication: SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.
Validation List no. 134 in IJSEM Online - Effective publication Online

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Pseudomonas oleovorans subsp. oleovorans (Lee and Chandler 1941) Saha et al. 2010, subsp. nov.
Type strain: (see also StrainInfo.net) ATCC 8062= CCUG 2087 = CFBP 5589 = CIP 59.11 = DSM 1045 = IFO (now NBRC) 13583 = JCM 11598 (IAM 1508) = LMG 2229 = NCIMB 6576 = NCTC 10692 = NRRL B-778 = VKM B-1522.
Sequence accession no. (16S rRNA gene) for the type strain: D84018.
Etymology: L. n. oleum, oil; L. part. adj. vorans, eating, devouring; N.L. part. adj. oleovorans, oil devouring.
Valid publication: Validation List no. 134. Int. J. Syst. Evol. Microbiol., 2010, 60, 1477-1479.
Effective publication: SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.
Validation List no. 134 in IJSEM Online - Effective publication Online

Notes:
1 The original authorship, Lee and Chandler 1941, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online
2 In the effective publication the strain NCIMB 6576 is erroneously cited as NCIB 6576.
Reference: ASSOCIATE EDITOR, IJSEM: Validation List no. 134 (footnote ¶). Int. J. Syst. Evol. Microbiol., 2010, 60, 1477-1479.
Validation List no. 134 in IJSEM Online

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Pseudomonas orientalis Dabboussi et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFML 96-170 = CFBP 4863 = CIP 105540 = DSM 17489.
Sequence accession no. (16S rRNA gene) for the type strain: AF064457.
Etymology: L. fem. adj. orientalis, Oriental, pertaining to the Orient.
Valid publication: Validation List no. 87. Int. J. Syst. Evol. Microbiol., 2002, 52, 1437-1438.
Effective publication: DABBOUSSI (F.), HAMZE (M.), ELOMARI (M.), VERHILLE (S.), BAIDA (N.), IZARD (D.) and LECLERC (H.): Taxonomic study of bacteria isolated from Lebanese spring waters: proposal of Pseudomonas cedrella sp. nov. and P. orientalis sp. nov. Res. Microbiol., 1999, 150, 303-316.
Validation List no. 87 in IJSEM Online - Effective publication Online

Note: At the time of publication, the type strain was not deposited in two publicly accessible service collections in different countries, and Pseudomonas orientalis Dabboussi et al. 2002 was not validly published [Rules 27(3) and 30]. However, according to the Judicial Opinion 81, Pseudomonas orientalis Dabboussi et al. 2002 is now considered to be validly published, since deposit in a second culture collection in a different country has been documented.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online

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Pseudomonas oryzihabitans Kodama et al. 1985, sp. nov.
Type strain: (see also StrainInfo.net) KS0036 = L-1 = AJ 2197 = ATCC 43272= CCUG 12540 = CIP 102996 = DSM 6835 = IAM 1568 = JCM 2952 = LMG 7040 = NBRC 102199.
Sequence accession no. (16S rRNA gene) for the type strain: D84004.
Etymology: L. n. oryza, rice; L. part. adj. habitans, inhabiting, dwelling; N.L. part. adj. oryzihabitans, rice-inhabiting.
Valid publication: KODAMA (K.), KIMURA (N.) and KOMAGATA (K.): Two new species of Pseudomonas: P. oryzihabitans isolated from rice paddy and clinical specimens and P. luteola isolated from clinical specimens. Int. J. Syst. Bacteriol., 1985, 35, 467-474.
Original article in IJSEM Online

Note: On page 473, the type strain is incorrectly cited as JCM 2592.

¤ Flavimonas oryzihabitans (Kodama et al. 1985) Holmes et al. 1987, comb. nov.

Pseudomonas oryzihabitans Kodama et al. 1985.
Type strain: (see also StrainInfo.net) KS0036 = L-1 = AJ 2197 = ATCC 43272= CCUG 12540 = CIP 102996 = DSM 6835 = IAM 1568 = JCM 2952.
Sequence accession no. (16S rRNA gene) for the type strain: D84004.
Reference: ANZAI (Y.), KUDO (Y.) and OYAIZU (H.): The phylogeny of the genera Chryseomonas, Flavimonas, and Pseudomonas supports synonymy of these three genera. Int. J. Syst. Bacteriol., 1997, 47, 249-251.
Original article in IJSEM Online

Note: The level of similarity between the 16S rRNA sequence of the strain KS0036 = L-1 = AJ 2197 = IAM 1568 = JCM 2952 (the type strain of the species Pseudomonas oryzihabitans Kodama et al. 1985) and the 16S rRNA of the strain DSM 50071 = ATCC 10145(the type strain of the species Pseudomonas aeruginosa (Schroeter 1872) Migula 1900 (Approved Lists 1980) is 93.9%. This result suggests that Flavimonas Holmes et al. 1987 is a later synonym of Pseudomonas Migula 1894 (Approved Lists 1980) and that the name Pseudomonas oryzihabitans should be used for this species.

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Pseudomonas otitidis Clark et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) MCC10330 = ATCC BAA-1130 = DSM 17224.
Sequence accession no. (16S rRNA gene) for the type strain: AY953147.
Etymology: Gr. n. ous otos, ear; N.L. suff. -itis -idis, used in names of inflammations; N.L. gen. n. otitidis, of inflammation of the ear.
Valid publication: CLARK (L.L.), DAJCS (J.J.), McLEAN (C.H.), BARTELL (J.G.) and STROMAN (D.W.): Pseudomonas otitidis sp. nov., isolated from patients with otic infections. Int. J. Syst. Evol. Microbiol., 2006, 56, 709-714.
Original article in IJSEM Online

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Pseudomonas pachastrellae Romanenko et al. 2005, sp. nov.
Type strain: (see also StrainInfo.net) CCUG 46540 = JCM 12285 = KMM 330 = NRIC 0583.
Sequence accession no. (16S rRNA gene) for the type strain: AB125366.
Etymology: N.L. gen. n. pachastrellae, of Pachastrella, the generic name of a sponge.
Valid publication: ROMANENKO (L.A.), UCHINO (M.), FALSEN (E.), FROLOVA (G.M.), ZHUKOVA (N.V.) and MIKHAILOV (V.V.): Pseudomonas pachastrellae sp. nov., isolated from a marine sponge. Int. J. Syst. Evol. Microbiol., 2005, 55, 919-924.
Original article in IJSEM Online

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Pseudomonas palleroniana Gardan et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFBP 4389 = ICMP 14253 = NCPPB 4278.
Sequence accession no. (16S rRNA gene) for the type strain: AY091527.
Etymology: N.L. fem. adj. palleroniana, pertaining to N. J. Palleroni, the famous microbiologist.
Valid publication: GARDAN (L.), BELLA (P.), MEYER (J.M.), CHRISTEN (R.), ROTT (P.), ACHOUAK (W.) and SAMSON (R.): Pseudomonas salomonii sp. nov., pathogenic on garlic, and Pseudomonas palleroniana sp. nov., isolated from rice. Int. J. Syst. Evol. Microbiol., 2002, 52, 2065-2074.
Original article in IJSEM Online

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Pseudomonas palleronii Davis 1970, species.
Type strain: (see also StrainInfo.net) Stanier 362t1 = ATCC 17724= CCUG 1780 = CCUG 20334 = CFBP 2445 = CIP 103304 = DSM 63 = JCM 21412 = LMG 2366 = NBRC 102513 = VKM B-1328.
Sequence accession no. (16S rRNA gene) for the type strain: AF078769.
Etymology: N.L. gen. masc. n. palleronii, of Palleroni, named after N.J. Palleroni, who first isolated this organism.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DAVIS (D.H.). In: DAVIS (D.H.), STANIER (R.Y.), DOUDOROFF (M.) and. MANDEL (M.): Taxonomic studies on some Gram-negative polarly flagellated "hydrogen bacteria" and related species. Archiv fur Mikrobiologie, 1970, 70, 1-13.

¤ Hydrogenophaga palleronii (Davis 1970) Willems et al. 1989, comb. nov.

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Pseudomonas panacis Park et al. 2005, sp. nov.
Type strain: (see also StrainInfo.net) CG20106 = CIP 108524 = IMSNU 14101 = KCTC 12330.
Sequence accession no. (16S rRNA gene) for the type strain: AY787208.
Etymology: L. gen. n. panacis, of panax, a fabulous plant supposed to heal all diseases, and the botanical genus name of ginseng (Panax L.).
Valid publication: PARK (Y.D.), LEE (H.B.), YI (H.), KIM (Y.), BAE (K.S.), CHOI (J.E.), JUNG (H.S.) and CHUN (J.): Pseudomonas panacis sp. nov., isolated from the surface of rusty roots of Korean ginseng. Int. J. Syst. Evol. Microbiol., 2005, 55, 1721-1724.
Original article in IJSEM Online

Note: In the paper by Park et al. 2005, the type strain IMSNU 14101 is erroneously cited as IMSNU 14100.

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Pseudomonas panipatensis Gupta et al. 2008, sp. nov.
Type strain: (see also StrainInfo.net) Esp-1 = CCM 7469 = MTCC 8990.
Sequence accession no. (16S rRNA gene) for the type strain: EF424401.
Etymology: N.L. fem. adj. panipatensis, pertaining to Panipat, the place of isolation of the bacterium.
Valid publication: GUPTA (S.K.), KUMARI (R.), PRAKASH (O.) and LAL (R.): Pseudomonas panipatensis sp. nov., isolated from an oil-contaminated site. Int. J. Syst. Evol. Microbiol., 2008, 58, 1339-1345.
Original article in IJSEM Online

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Pseudomonas parafulva Uchino et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CB-1 = AJ 2129 = CIP 107617 = DSM 17004 = IFO (now NBRC) 16636 = JCM 11244 = NRIC 0501.
Sequence accession no. (16S rRNA gene) for the type strain: AB060132.
Etymology: Gr. prep. para, beside, alongside of, near, like; L. fem. adj. fulva, a specific epithet; N.L. fem. adj. parafulva, (Pseudomonas) fulva-like.
Valid publication: Validation List no. 85. Int. J. Syst. Evol. Microbiol., 2002, 52, 685-690.
Effective publication: UCHINO (M.), SHIDA (O.), UCHIMURA (T.) and KOMAGATA (K.): Recharacterization of Pseudomonas fulva Iizuka and Komagata 1963, and proposals of Pseudomonas parafulva sp. nov. and Pseudomonas cremoricolorata sp. nov. J. Gen. Appl. Microbiol., 2001, 47, 247-261.
Validation List no. 85 in IJSEM Online - Effective publication Online

Notes:
1 In Validation List no. 85, the reference of the effective publication is erroneously cited as J. Gen. Appl. Microbiol., 2001, 46, 247-261.
2 According to Rules 27(3) and 30, this name is not validly published, because the original publication makes reference to the deposit of the type strain in a single recognized culture collection. However, according to the Judicial Opinion 81, Pseudomonas parafulva Uchino et al. 2002 is considered to be validly published, since deposit in a second culture collection in a different country has been documented.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online

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Pseudomonas paucimobilis Holmes et al. 1977, species.
Type strain: (see also StrainInfo.net) ATCC 29837= CCUG 6518 = CCUG 31192 = CIP 100752 = DSM 1098 = GIFU 2395 = HAMBI 1744 = IFO (now NBRC) 13935 = JCM 7516 = LMG 1227 = NCAIM B.01654 = NCTC 11030 = VKM B-2241.
Sequence accession no. (16S rRNA gene) for the type strain: AM237364.
Etymology: L. adj. paucus, few, little; L. adj. mobilis, movable, mobile; N.L. fem. adj. paucimobilis, intended to mean few motile cells.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HOLMES (B.), OWEN (R.J.), EVANS (A.), MALNICK (H.) and WILLCOX (W.R.): Pseudomonas paucimobilis, a new species isolated from human clinical specimens, the hospital environment, and other sources. International Journal of Systematic Bacteriology, 1977, 27, 133-146.

¤ Sphingomonas paucimobilis (Holmes et al. 1977) Yabuuchi et al. 1990, comb. nov.

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Pseudomonas pelagia Hwang et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) CL-AP6 = JCM 15562 = KCCM 90073.
Sequence accession no. (16S rRNA gene) for the type strain: EU888911.
Etymology: L. fem. adj. pelagia, of or belonging to the sea.
Valid publication: HWANG (C.Y.), ZHANG (G.I.), KANG (S.H.), KIM (H.J.) and CHO (B.C.): Pseudomonas pelagia sp. nov., isolated from a culture of the Antarctic green alga Pyramimonas gelidicola. Int. J. Syst. Evol. Microbiol., 2009, 59, 3019-3024.
Original article in IJSEM Online

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Pseudomonas peli Vanparys et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) R-20805 = DSM 17833 = LMG 23201.
Sequence accession no. (16S rRNA gene) for the type strain: AM114534.
Etymology: Gr. n. pelos, sludge; N.L. gen. n. peli, of sludge.
Valid publication: VANPARYS (B.), HEYLEN (K.), LEBBE (L.) and DE VOS (P.): Pseudomonas peli sp. nov. and Pseudomonas borbori sp. nov., isolated from a nitrifying inoculum. Int. J. Syst. Evol. Microbiol., 2006, 56, 1875-1881.
Original article in IJSEM Online

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Pseudomonas perfectomarina (ex ZoBell and Upham 1944) Baumann et al. 1983, sp. nov., nom. rev.
Type strain: (see also StrainInfo.net) 218 = ATCC 14405.
Sequence accession no. (16S rRNA gene) for the type strain: U65012.
Synonym: "Pseudomonas perfectomarinus" (sic) ZoBell and Upham 1944.
Etymology: L. part. adj. perfectus, complete, perfect; L. fem. adj. marina, of the sea, marine; N.L. fem. adj. perfectomarina, completely marine.
Valid publication: BAUMANN (P.), BOWDITCH (R.D.), BAUMANN (L.) and BEAMAN (B.): Taxonomy of marine Pseudomonas species: P. stanieri sp. nov.; P. perfectomarina sp. nov., nom. rev.; P. nautica; and P. doudoroffii. Int. J. Syst. Bacteriol., 1983, 33, 857-865.
Original article in IJSEM Online

¤ Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980).

Notes:
1 According to Döhler et al. 1987, Pseudomonas perfectomarina (ex ZoBell and Upham 1944) Baumann et al. 1983 is a later heterotypic synonym of ¤ Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980).
Reference: DÖHLER (K.), HUSS (V.A.R.) and ZUMFT (W.G.): Transfer of Pseudomonas perfectomarina Baumann, Bowditch, Baumann, and Beaman 1983 to Pseudomonas stutzeri (Lehmann and Neumann 1896) Sidjerius 1946. Int. J. Syst. Bacteriol., 1987, 37, 1-3.
Original article in IJSEM Online
2 Rosselló-Mora et al. clarified the taxonomic position of Pseudomonas perfectomarina in the species Pseudomonas stutzeri and concluded that the strain ATCC 14405belongs to genomovar 2 of Pseudomonas stutzeri.
Reference: ROSSELLÓ-MORA (R.A.), GARCÍA-VALDÉS (E.) and LALUCAT (J.): Taxonomic relationship between Pseudomonas perfectomarina ZoBell and Pseudomonas stutzeri. Int. J. Syst. Bacteriol., 1993, 43, 852-854.
Original article in IJSEM Online

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Pseudomonas pertucinogena Kawai and Yabuuchi 1975, species.
Type strain: (see also StrainInfo.net) ATCC 190= CCUG 7832 = CIP 106696 = IFO (now NBRC) 14163 = JCM 11590 = LMG 1874.
Sequence accession no. (16S rRNA gene) for the type strain: AB021380.
Etymology: N.L. n. pertucinum, pertucin, a bacteriocin active against Bordetella pertussis; L. suff. genus -a -um (from L. v. gigno, to produce, give birth to, beget), producing; N.L. fem. adj. pertucinogena, intended to mean pertucin producing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: KAWAI (Y.) and YABUUCHI (E.): Pseudomonas pertucinogena sp. n., an organism previously misidentified as Bordetella pertussis. International Journal of Systematic Bacteriology, 1975, 25, 317-323.

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Pseudomonas phenazinium Bell and Turner 1973, species.
Type strain: (see also StrainInfo.net) ATCC 33666= CCUG 20836 = CFBP 4793 = CIP 106502 = DSM 10684 = JCM 10564 = LMG 2247 = NCIB (now NCIMB) 11027.
Sequence accession no. (16S rRNA gene) for the type strain: U96936.
Etymology: Orthography and etymology uncertain; possibly refers to iodinin, which is a phenazine pigment.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: BELL (S.C.) and TURNER (J.M.): Iodinin biosynthesis by a pseudomonad. Biochemical Society Transactions, 1973, 1, 751-753.

¤ Burkholderia phenazinium (Bell and Turner 1973) Viallard et al. 1998, comb. nov.

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Pseudomonas pickettii Ralston et al. 1973, species.
Type strain: (see also StrainInfo.net) ATCC 27511= CCUG 3318 = CFBP 2459 = CIP 73.23 = DSM 6297 = HAMBI 2158 = JCM 5969 = LMG 5942 = NCTC 11149.
Sequence accession no. (16S rRNA gene) for the type strain: AY741342.
Etymology: N.L. gen. masc. n. pickettii, of Pickett; named after M.J. Pickett.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: RALSTON (E.), PALLERONI (N.J.) and DODOROFF (M.): Pseudomonas pickettii, a new species of clinical origin related to Pseudomonas solanacearum. International Journal of Systematic Bacteriology, 1973, 23, 15-19.

¤ Burkholderia pickettii (Ralston et al. 1973) Yabuuchi et al. 1993, comb. nov.

¤ Ralstonia pickettii (Ralston et al. 1973) Yabuuchi et al. 1996, comb. nov.

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Pseudomonas pictorum Gray and Thornton 1928, species.
Type strain: (see also StrainInfo.net) ATCC 23328= CCUG 1823 = CCUG 3368 = CIP 103273 = JCM 9942 = LMG 981 = NRRL B-2543 = VKM B-1240.
Sequence accession no. (16S rRNA gene) for the type strain: AJ131116.
Etymology: L. gen. pl. n. pictorum, of painters; here, intended to mean of the Picts, named after the Picts, a scottish tribe.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: GRAY (P.H.H.) and THORNTON (H.G.): Soil bacteria that decompose certain aromatic compounds. Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung II, 1928, 73, 74-96.

Note: According to Anzai et al., Pseudomonas pictorum Gray and Thornton 1928 (Approved Lists 1980) does not belong to the genus Pseudomonas sensu stricto.
Reference: ANZAI (Y.), KIM (H.), PARK (J.Y.), WAKABAYASHI (H.) and OYAIZU (H.): Phylogenetic affiliation of the pseudomonads based on 16S rRNA sequence. Int. J. Syst. Evol. Microbiol., 2000, 50, 1563-1589.

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Pseudomonas plantarii Azegami et al. 1987, sp. nov.
Type strain: (see also StrainInfo.net) AZ 8201 = ATCC 43733= CFBP 3573 = CFBP 3997 = CIP 105769 = DSM 9509 = ICMP 9424 = JCM 5492 = LMG 9035 = NCPPB 3590 = NIAES 1723.
Sequence accession no. (16S rRNA gene) for the type strain: U96933.
Etymology: L. n. plantarium, nursery, seedbed; L. gen. n. plantarii, of seedbed.
Valid publication: AZEGAMI (K.), NISHIYAMA (K.), WATANABE (Y.), KADOTA (I.), OHUCHI (A.) and FUKAZAWA (C.): Pseudomonas plantarii sp. nov., the causal agent of rice seedling blight. Int. J. Syst. Bacteriol., 1987, 37, 144-152.
Original article in IJSEM Online

¤ Burkholderia plantarii (Azegami et al. 1987) Urakami et al. 1994, comb. nov.

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Pseudomonas plecoglossicida Nishimori et al. 2000, sp. nov.
Type strain: (see also StrainInfo.net) FPC 951 = ATCC 700383= CCUG 49623 = CIP 106493 = DSM 15088 = JCM 21584 = NBRC 103162.
Sequence accession no. (16S rRNA gene) for the type strain: AB009457.
Etymology: N.L. n. Plecoglossus, systematic name of afish genus; L. suff. -cida (from L. v. caedere, to kill), killer; N.L. n. plecoglossicida, the killer of Plecoglossus, the genus to which ayu belongs.
Valid publication: NISHIMORI (E.), KITA-TSUKAMOTO (K.) and WAKABAYASHI (H.): Pseudomonas plecoglossicida sp. nov., the causative agent of bacterial haemorrhagic ascites of ayu, Plecoglossus altivelis. Int. J. Syst. Evol. Microbiol., 2000, 50, 83-89.
Original article in IJSEM Online

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Pseudomonas poae Behrendt et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) P 527/13 = DSM 14936 = LMG 21465.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492829.
Etymology: Gr. n. poa, grass; N.L. gen. n. poae of/from grass.
Valid publication: BEHRENDT (U.), ULRICH (A.) and SCHUMANN (P.): Fluorescent pseudomonads associated with the phyllosphere of grasses; Pseudomonas trivialis sp. nov., Pseudomonas poae sp. nov. and Pseudomonas congelans sp. nov. Int. J. Syst. Evol. Microbiol., 2003, 53, 1461-1469.
Original article in IJSEM Online

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Pseudomonas pohangensis Weon et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) H3-R18 = DSM 17875 = KACC 11517.
Sequence accession no. (16S rRNA gene) for the type strain: DQ339144.
Etymology: N.L. fem. adj. pohangensis, referring to Pohang city in Korea, where the type strain was found.
Valid publication: WEON (H.Y.), KIM (B.Y.), YOO (S.H.), BAEK (Y.K.), LEE (S.Y.), KWON (S.W.), GO (S.J.) and STACKEBRANDT (E.): Pseudomonas pohangensis sp. nov., isolated from seashore sand in Korea. Int. J. Syst. Evol. Microbiol., 2006, 56, 2153-2156.
Original article in IJSEM Online

Pseudomonas pohangensis Weon et al. 2006 emend. Hameed et al. 2014.
Reference: HAMEED, A., SHAHINA, M., LIN, S.-Y., LIU, Y.-C. and YOUNG, C.-C. 2014. Pseudomonas hussainii sp. nov., isolated from droppings of a seashore bird, and emended descriptions of Pseudomonas pohangensis, Pseudomonas benzenivorans and Pseudomonas segetis. Int. J. Syst. Evol. Microbiol., 64, 2330-2337.
Original article.

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Pseudomonas populi Anwar et al. 2016, sp. nov.
Type strain: (see also StrainInfo.net) KBL-4-9=JCM 19138=CCTCC AB 2013069=NRRL B-59988.
Sequence accession no. (16S rRNA) for the type strain: KC609749.
Whole-genome sequence accession no. for the type strain: -.
Etymology: L. fem. gen. n. populi, of a poplar tree, pertaining to Populus euphratica, the Latin name for the poplars that grow in the forest from which the type strain was isolated.
Source: Plant.
Valid publication: ANWAR, N., ABAYDULLA, G., ZAYADAN, B., ABDURAHMAN, M., HAMOOD, B., ERKIN, R., ISMAYIL, N., ROZAHON, M., MAMTIMIN, H. and RAHMAN, E. 2016. Pseudomonas populi sp. nov., an endophytic bacterium isolated from Populus euphratica. Int. J. Syst. Evol. Microbiol., 66, 1419-1425.
Original article in IJSEM Online

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Pseudomonas prosekii Kosina et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) AN/28/1=CCM 7990=LMG 26867.
Sequence accession no. (16S rRNA) for the type strain: JN814372.
Whole-genome sequence accession no. for the type strain: -.
Etymology: -.
Source: Environmental.
Valid publication: Validation List no. 158. List of new names and new combinations previously effectively, but not validly, published. Int. J Syst. Evol. Microbiol., 64 (2014), 2184-2187.
Validation List Online
Effective publication: KOSINA, M., BARTÁK, M., MAŠLAŇOVÁ, I., PASCUTTI, A. V., SEDO, O., LEXA, M. and SEDLÁ ČEK, I. 2013. Pseudomonas prosekii sp. nov., a novel psychrotrophic bacterium from Antarctica. Curr. Microbiol., 67, 637-646.
Effective publication Online

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Pseudomonas protegens Ramette et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) CHA0 = CFBP 6595 = DSM 19095.
Sequence accession no. (16S rRNA gene) for the type strain: AJ278812.
Etymology: L. part. adj. protegens, protecting, referring to the ability of the organism to protect plants from soil-borne phytopathogens.
Valid publication: Validation List no. 146. Int. J. Syst. Evol. Microbiol., 2012, 62, 1443-1445.
Effective publication: RAMETTE (A.), FRAPOLLI (M.), FISCHER-LE SAUX (M.), GRUFFAZ (C.), MEYER (J.M.), DÉFAGO (G.), SUTRA (L.) and MOËNNE-LOCCOZ (Y.): Pseudomonas protegens sp. nov., widespread plant-protecting bacteria producing the biocontrol compounds 2,4-diacetylphloroglucinol and pyoluteorin. Syst. Appl. Microbiol., 2011, 34, 180-188.
Validation List no. 146 in IJSEM Online - Effective publication Online

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Pseudomonas proteolytica Reddy et al. 2004, sp. nov.
Type strain: (see also StrainInfo.net) CMS 64 = CCUG 51515 = DSM 15321 = MTCC 4994.
Sequence accession no. (16S rRNA gene) for the type strain: AJ537603.
Etymology: N.L. fem. adj. proteolytica, proteolytic.
Valid publication: REDDY (G.S.N.), MATSUMOTO (G.I.), SCHUMANN (P.), STACKEBRANDT (E.) and SHIVAJI (S.): Psychrophilic pseudomonads from Antarctica: Pseudomonas antarctica sp. nov., Pseudomonas meridiana sp. nov. and Pseudomonas proteolytica sp. nov. Int. J. Syst. Evol. Microbiol., 2004, 54, 713-719.
Original article in IJSEM Online

Pseudomonas proteolytica Reddy et al. 2004 emend. van den Beld et al. 2016.
Reference: VAN DEN BELD, M. J., REINDERS, E., NOTERMANS, D. W. and REUBSAET, F. A. 2016. Possible misidentification of species in the Pseudomonas fluorescens lineage as Burkholderia pseudomallei and Francisella tularensis, and emended descriptions of Pseudomonas brenneri, Pseudomonas gessardii and Pseudomonas proteolytica. Int. J. Syst. Evol. Microbiol., 66, 3420-3425.
Original article.

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Pseudomonas pseudoalcaligenes Stanier 1966, species.
Type strain: (see also StrainInfo.net) ATCC 17440= CCUG 51525 = CFBP 2435 = CIP 66.14 = DSM 50188 = IFO (now NBRC) 14167 = JCM 5968 = LMG 1225 = NCCB 76045 = NCTC 10860.
Sequence accession no. (16S rRNA gene) for the type strain: Z76666.
Etymology: Gr. adj.  pseudês, false; N.L. n. alcali (from Arabic article al, the; Arabic n. qaliy, ashes of saltwort), alkali; N.L. suff. -genes (from Gr. v. gennaô, to produce), producing; N.L. adj. alcaligenes, alkali-producing; N.L. adj. pseudoalcaligenes, false alkaliproducing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: STANIER (R.Y.). In: STANIER (R.Y.), PALLERONI (N.J.) and DOUDOROFF (M.): The aerobic pseudomonads: a taxonomic study. Journal of General Microbiology, 1966, 43, 159-271.

¤ Pseudomonas oleovorans Lee and Chandler 1941 (Approved Lists 1980).

Note: According to Saha et al. 2010, Pseudomonas pseudoalcaligenes Stanier 1966 (Approved Lists 1980) is a later heterotypic synonym of ¤ Pseudomonas oleovorans Lee and Chandler 1941 (Approved Lists 1980).
References: List of Changes in Taxonomic Opinion no. 12. Int. J. Syst. Evol. Microbiol., 2010, 60, 1482-1483 [SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.]
List of Changes in Taxonomic Opinion no. 12 in IJSEM Online - Effective publication Online

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Pseudomonas pseudoalcaligenes subsp. citrulli Schaad et al. 1978, subspecies.
Type strain: (see also StrainInfo.net) FC-247 = ATCC 29625= CCUG 17393 = CFBP 4459 = CIP 106436 = IBSBF 1851 = ICMP 7500 = ICPB 30064 = LMG 5376 = NCPPB 3679.
Sequence accession no. (16S rRNA gene) for the type strain: AF137506.
Etymology: N.L. n. Citrullus, a genus of melon plants; N.L. gen. n. citrulli, of Citrullus.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: SCHAAD (N.W.), SOWELL (G.), GOTH (R.W.), COLWELL (R.R.) and WEBB (R.E.): Pseudomonas pseudoalcaligenes subsp. citrulli subsp. nov. International Journal of Systematic Bacteriology, 1978, 28, 117-125.

¤ Pseudomonas avenae subsp. citrulli (Schaad et al. 1978) Hu et al. 1991, comb. nov.

¤ Acidovorax avenae subsp. citrulli (Schaad et al. 1978) Willems et al. 1992, comb. nov.

¤ Acidovorax citrulli (Schaad et al. 1978) Schaad et al. 2009, comb. nov.

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Pseudomonas pseudoalcaligenes subsp. konjaci Goto 1983, subsp. nov.
Type strain: (see also StrainInfo.net) K2 = ATCC 33996= CCUG 17394 = CFBP 4460 = CIP 106439 = DSM 7481 = ICMP 7733 = JCM 2397 = LMG 5691 = NCPPB 3698 = PDDCC 7733.
Sequence accession no. (16S rRNA gene) for the type strain: AJ420325.
Etymology: N.L. gen. n. konjaci, of the konjac plant, Amorphophallus rivieri cv. Konjac.
Valid publication: GOTO (M.): Pseudomonas pseudoalcaligenes subsp. konjaci subsp. nov., the causal agent of bacterial leaf blight of konjac (Amorphophalus konjac Koch.). Int. J. Syst. Bacteriol., 1983, 33, 539-545.
Original article in IJSEM Online

¤ Pseudomonas avenae subsp. konjaci (Goto 1983) Hu et al. 1991, comb. nov.

¤ Acidovorax konjaci (Goto 1983) Willems et al. 1992, comb. nov.

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Pseudomonas pseudoalcaligenes subsp. pseudoalcaligenes (Stanier 1966) Schaad et al. 1978, subspecies.
Type strain: (see also StrainInfo.net) ATCC 17440= CCUG 51525 = CFBP 2435 = CIP 66.14 = DSM 50188 = IFO (now NBRC) 14167 = JCM 5968 = LMG 1225 = NCCB 76045 = NCTC 10860.
Sequence accession no. (16S rRNA gene) for the type strain: Z76666.
Etymology: Gr. adj.  pseudês, false; N.L. n. alcali (from Arabic article al, the; Arabic n. qaliy, ashes of saltwort), alkali; N.L. suff. -genes (from Gr. v. gennaô, to produce), producing; N.L. adj. alcaligenes, alkali-producing; N.L. adj. pseudoalcaligenes, false alkaliproducing.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: STANIER (R.Y.). In: STANIER (R.Y.), PALLERONI (N.J.) and DOUDOROFF (M.): The aerobic pseudomonads: a taxonomic study. Journal of General Microbiology, 1966, 43, 159-271.

Notes:
1 The original authorship, Stanier 1966, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online
2 If an author accepts that Pseudomonas pseudoalcaligenes subsp. citrulli Schaad et al. 1978 (Approved Lists 1980) and Pseudomonas pseudoalcaligenes subsp. konjaci Goto 1983 are subspecies of Pseudomonas avenae Manns 1909 (Approved Lists 1980), then Pseudomonas pseudoalcaligenes subsp. pseudoalcaligenes Stanier 1966 (Approved Lists 1980) should thereafter be designated simply as Pseudomonas pseudoalcaligenes Stanier 1966 (Approved Lists 1980).

¤ Pseudomonas oleovorans Lee and Chandler 1941 (Approved Lists 1980).

Note: According to Saha et al. 2010, Pseudomonas pseudoalcaligenes Stanier 1966 (Approved Lists 1980) is a later heterotypic synonym of ¤ Pseudomonas oleovorans Lee and Chandler 1941 (Approved Lists 1980).
References: List of Changes in Taxonomic Opinion no. 12. Int. J. Syst. Evol. Microbiol., 2010, 60, 1482-1483 [SAHA (R.), SPRÖER (C.), BECK (B.) and BAGLEY (S.): Pseudomonas oleovorans subsp. lubricantis subsp. nov., and reclassification of Pseudomonas pseudoalcaligenes ATCC 17440T as later synonym of Pseudomonas oleovorans ATCC 8062T. Curr. Microbiol., 2010, 60, 294-300.]
List of Changes in Taxonomic Opinion no. 12 in IJSEM Online - Effective publication Online

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Pseudomonas pseudoflava Auling et al. 1978, species.
Type strain: (see also StrainInfo.net) GA3 = ATCC 33668= CCUG 13799 = CFBP 2439 = CIP 103270 = DSM 1034 = JCM 21410 = LMG 5945 = NBRC 102511.
Sequence accession no. (16S rRNA gene) for the type strain: AF078770.
Etymology: Gr. adj. pseudês, false; L. fem. adj. flava, yellow, and also a specific epithet; N.L. fem. adj. pseudoflava, not the true (Pseudomonas) flava, referring to the close relationship to Pseudomonas flava.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: AULING (G.), REH (M.), LEE (C.M.) and SCHLEGEL (H.G.): Pseudomonas pseudoflava, a new species of hydrogen-oxidizing bacteria: its differentiation from Pseudomonas flava and other yellow-pigmented, Gram-negative, hydrogen oxidizing species. International Journal of Systematic Bacteriology, 1978, 28, 82-95.

¤ Hydrogenophaga pseudoflava (Auling et al. 1978) Willems et al. 1989, comb. nov.

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Pseudomonas pseudomallei (Whitmore 1913) Haynes 1957, species.
Type strain: (see also StrainInfo.net) WRAIR 286 = ATCC 23343= NCTC 12939.
Sequence accession no. (16S rRNA gene) for the type strain: DQ108392.
Synonyms: "Bacillus pseudomallei" Whitmore 1913, "Bacterium whitmori" Stanton and Fletcher 1921, "Malleomyces pseudomallei" Breed 1939, "Loefflerella pseudomallei" Brindle and Cowan 1951.
Etymology: Gr. adj.  pseudês, false; L. n. malleus, a hammer, maul, the disease of glanders, and also a specific epithet; N.L. gen. n. pseudomallei, of false glanders, the false (Pseudomonas) mallei.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 WHITMORE (A.): An account of a glanders-like disease occurring in Rangoon. Journal of Hygiene, 1913, 13, 1-34. 2 HAYNES (W.C.): Genus I. Pseudomonas Migula 1894. In: R.S. BREED, E.G.D. MURRAY and N.R. SMITH (editors): Bergey's Manual of Determinative Bacteriology, 7th edition, The Williams & Wilkins Co, Baltimore, 1957, pp. 89-152.

¤ Burkholderia pseudomallei (Whitmore 1913) Yabuuchi et al. 1993, comb. nov.

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Pseudomonas psychrophila Yumoto et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) E-3 = CIP 107901 = JCM 10889.
Sequence accession no. (16S rRNA gene) for the type strain: AB041885.
Etymology: Gr. adj. psuchros, cold; N.L. adj. philus -a -um (from Gr. adj. philos -ê -on), friend, loving; N.L. fem. adj. psychrophila, cold loving.
Valid publication: Validation List no. 85. Int. J. Syst. Evol. Microbiol., 2002, 52, 685-690.
Effective publication: YUMOTO (I.), KUSANO (T.), SHINGYO (T.), NODASAKA (Y.), MATSUYAMA (H.) and OKUYAMA (H.): Assignment of Pseudomonas sp. strain E-3 to Pseudomonas psychrophila sp. nov., a new facultatively psychrophilic bacterium. Extremophiles, 2001, 5, 343-349.
Validation List no. 85 in IJSEM Online - Effective publication Online

Note: According to Rules 27(3) and 30, this name is not validly published, because the original publication makes reference to the deposit of the type strain in a single recognized culture collection. However, according to the Judicial Opinion 81, Pseudomonas psychrophila Yumoto et al. 2002 is considered to be validly published, since deposit in a second culture collection in a different country has been documented.
References:
1 EUZÉBY (J.P.) and TINDALL (B.J.): Status of strains that contravene Rules 27(3) and 30 of the Bacteriological Code. Request for an Opinion. Int. J. Syst. Evol. Microbiol., 2004, 54, 293-301.
Original article in IJSEM Online
2 JUDICIAL COMMISSION OF THE INTERNATIONAL COMMITTEE ON SYSTEMATICS OF PROKARYOTES: Status of strains that contravene Rules 27 (3) and 30 of the International Code of Nomenclature of Bacteria. Opinion 81. Int. J. Syst. Evol. Microbiol., 2008, 58, 1755-1763.
Opinion 81 in IJSEM Online

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Pseudomonas psychrotolerans Hauser et al. 2004, sp. nov.
Type strain: (see also StrainInfo.net) C36 = CCUG 51516 = DSM 15758 = LMG 21977.
Sequence accession no. (16S rRNA gene) for the type strain: AJ575816.
Etymology: Gr. adj. psuchros, cold; L. part. adj. tolerans, tolerating; N.L. part. adj. psychrotolerans, cold-tolerating.
Valid publication: HAUSER (E.), KÄMPFER (P.) and BUSSE (H.J.): Pseudomonas psychrotolerans sp. nov. Int. J. Syst. Evol. Microbiol., 2004, 54, 1633-1637.
Original article in IJSEM Online

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Pseudomonas punonensis Ramos et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) LMT03 = CECT 8089 = LMG 26839.
Sequence accession no. (16S rRNA gene) for the type strain: JQ344321.
Etymology: N.L. fem. adj. punonensis, of or belonging to Puno, a region of Peru where the type strain was isolated.
Valid publication: RAMOS (E.), RAMÍREZ-BAHENA (M.H.), VALVERDE (A.), VELÁZQUEZ (E.), ZÚÑIGA (D.), VELEZMORO (C.) and PEIX (A.): Pseudomonas punonensis sp. nov., isolated from straw. Int. J. Syst. Evol. Microbiol., 2013, 63, 1834-1839.
Original article in IJSEM Online

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Pseudomonas putida (Trevisan 1889) Migula 1895, species.
Type strain: (see also StrainInfo.net) ATCC 12633= CCUG 12690 = CFBP 2066 = CIP 52.191 = DSM 291 = HAMBI 7 = IFO (now NBRC) 14164 = JCM 13063 = JCM 20120 = LMG 2257 = NCAIM B.01634 = NCCB 72006 = NCCB 68020 = NCTC 10936.
Sequence accession no. (16S rRNA gene) for the type strain: D84020.
Synonyms: "Bacillus fluorescens putidus" Flügge 1886, "Bacillus putidus" Trevisan 1889, "Pseudomonas eisenbergii" Migula 1900, "Pseudomonas convexa" Chester 1901, "Pseudomonas incognita" Chester 1901, "Pseudomonas ovalis" Chester 1901, "Pseudomonas rugosa" (Wright 1895) Chester 1901, "Pseudomonas striata" Chester 1901.
Etymology: L. fem. adj. putida, stinking, fetid.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 TREVISAN (V.): I. Generi e le Specie delle Batteriacee, Zanaboni and Gabuzzi, Milano, 1889. 2 MIGULA (W.): Schizomycetes (Bacteria, Bacterien). In: A. ENGLER and K. PRANTL (editors): Die näturlichen Pfanzenfamilien, W. Engelmann, Leipzig, Teil I, Abteilung Ia, 1895, pp. 1-44.

Note: Pseudomonas putida (Trevisan 1889) Migula 1895 (Approved Lists 1980) is an earlier synonym of ¤ Arthrobacter siderocapsulatus Dubinina and Zhdanov 1975 (Approved Lists 1980).
Reference: CHUN (J.), RHEE (M.S.), HAN (J.I.) and BAE (K.S.): Arthrobacter siderocapsulatus Dubinina and Zhdanov 1975AL is a later subjective synonym of Pseudomonas putida (Trevisan 1889) Migula 1895AL. Int. J. Syst. Evol. Microbiol., 2001, 51, 169-170.
Original article in IJSEM Online

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Pseudomonas pyrrocinia Imanaka et al. 1965, species.
Type strain: (see also StrainInfo.net) ATCC 15958= CFBP 4794 = CIP 105874 = DSM 10685 = LMG 14191.
Sequence accession no. (16S rRNA gene) for the type strain: U96930.
Etymology: Etymology uncertain, possibly N.L. adj. pyrrocinia, referring to the antibiotic properties of pyrrolnitrin, which is produced by strains of this species.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IMANAKA (H.), KOUSAKA (M.), TAMURA (G.) and ARIMA (K.): Studies on pyrrolnitrin, a new antibiotic. II. Taxonomic studies on pyrrolnitrin-producing strain. Journal of Antibiotics (Tokyo) Series A, 1965, 18, 205-206.

¤ Burkholderia pyrrocinia (Imanaka et al. 1965) Vandamme et al. 1997, comb. nov.

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Pseudomonas radiora Ito and Iizuka 1971, species.
Type strain: (see also StrainInfo.net) ATCC 27329= CIP 101128 = DSM 1819 = IFO (now NBRC) 15690 = JCM 2831 = LMG 2269 = NCIB (now NCIMB) 10815 = VKM B-2144.
Sequence accession no. (16S rRNA gene) for the type strain: D32227, CP001001 (complete genome).
Etymology: The etymology is uncertain. Perhaps, intended to mean pertaining to radiation.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: ITO (H.) and IIZUKA (H.): Taxonomic studies on radio-resistant Pseudomonas. Part XII. Studies on the microorganisms of cereal grains. Agricultural and Biological Chemistry, 1971, 35, 1566-1571.

Note: The specific epithet radiora (sic) has been corrected in the new combination Methylobacterium radiotolerans [see, ¤ Methylobacterium radiotolerans corrig. (Ito and Iizuka 1971) Green and Bousfield 1983, comb. nov.]. However the name "Pseudomonas radiotolerans" corrig. has never been formally proposed before December 14, 2000 (date of publication of the amended Rule 61; see Int. J. Syst. Evol. Microbiol., 2000, 50, page 2239, column 2). Consequently, in the "List of Prokaryotic names with Standing in Nomenclature", the epithet radiora (sic) has been maintained for Pseudomonas radiora.

¤ Methylobacterium radiotolerans corrig. (Ito and Iizuka 1971) Green and Bousfield 1983, comb. nov.

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Pseudomonas reinekei Cámara et al. 2007, sp. nov.
Type strain: (see also StrainInfo.net) MT1 = CCUG 53116 = DSM 18361.
Sequence accession no. (16S rRNA gene) for the type strain: AM293565.
Etymology: N.L. gen. masc. n. reinekei, of Reineke, named in honour of Walter Reineke, for his pioneering contributions to the study of the biodegradation of chlorinated aromatics.
Valid publication: CÁMARA (B.), STRÖMPL (C.), VERBARG (S.), SPRÖER (C.), PIEPER (D.H.) and TINDALL (B.J.): Pseudomonas reinekei sp. nov., Pseudomonas moorei sp. nov. and Pseudomonas mohnii sp. nov., novel species capable of degrading chlorosalicylates or isopimaric acid. Int. J. Syst. Evol. Microbiol., 2007, 57, 923-931.
Original article in IJSEM Online

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Pseudomonas resinovorans Delaporte et al. 1961, species.
Type strain: (see also StrainInfo.net) ATCC 14235= CCUG 2473 = CCUG 4439 = CFBP 5590 = CIP 61.9 = LMG 2274 = NRRL B-2649.
Sequence accession no. (16S rRNA gene) for the type strain: Z76668.
Etymology: L. n. resina, resin; L. part. adj. vorans, eating, devouring; N.L. part. adj. resinovorans, resin devouring.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DELAPORTE (B.), RAYMOND (M.) and DASTE (P.): Une bactérie du sol capable d'utiliser, comme source de carbone, la fraction fixe de certaines oléorésines, Pseudomonas resinovorans n. sp. Compte Rendu des Séances de l'Académie des Sciences, Paris, 1961, 252, 1073-1075.

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Pseudomonas rhizosphaerae Peix et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) IH5 = CECT 5726 = LMG 21640.
Sequence accession no. (16S rRNA gene) for the type strain: AY152673.
Etymology: Gr. fem. n. rhiza, root; L. fem. n. sphaera (from Gr. fem. n. sphaira), ball, globe, sphere; N.L. gen. fem. n. rhizosphaerae, of the rhizosphere.
Valid publication: PEIX (A.), RIVAS (R.), MATEOS (P.F.), MARTÍNEZ-MOLINA (E.), RODRÍGUEZ-BARRUECO (C.) and VELÁZQUEZ (E.): Pseudomonas rhizosphaerae sp. nov., a novel species that actively solubilizes phosphate in vitro. Int. J. Syst. Evol. Microbiol., 2003, 53, 2067-2072.
Original article in IJSEM Online

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Pseudomonas rhodesiae Coroler et al. 1997, sp. nov.
Type strain: (see also StrainInfo.net) CCUG 38732 = CIP 104664 = DSM 14020 = JCM 11940 = LMG 17764.
Sequence accession no. (16S rRNA gene) for the type strain: AF064459.
Etymology: N.L. fem. gen. n. rhodesiae, of Rhodes, in honor of M.R. Rhodes, an English microbiologist.
Valid publication: Validation List no. 61. Int. J. Syst. Bacteriol., 1997, 47, 601-602.
Effective publication: COROLER (L.), ELOMARI (M.), HOSTE (B.), GILLIS (M.), IZARD (D.) and LECLERC (H.): Pseudomonas rhodesiae sp. nov., a new species isolated from natural mineral waters. Syst. Appl. Microbiol., 1996, 19, 600-607.
Validation List no. 61 in IJSEM Online - Effective publication Online

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Pseudomonas rhodos Heumann 1962, species.
Type strain: (see also StrainInfo.net) ATCC 14821= CIP 101127 = DSM 2163 = IFO (now NBRC) 15691 = JCM 2811 = LMG 2275 = NCIB (now NCIMB) 9421 = VKM B-2065.
Sequence accession no. (16S rRNA gene) for the type strain: AB175644.
Etymology: Gr. n. rhodon, the rose; N.L. fem. adj. rhodos (sic), rose colored.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HEUMANN (W.): Die Methodik der Kreuzung Sternbildender Bakterien. Biolgisches Zentralblatt, 1962, 81, 341-354.

Note: The specific epithet rhodos (sic) has been corrected in the new combination Methylobacterium rhodinum [see, ¤ Methylobacterium rhodinum corrig. (Heumann 1962) Green and Bousfield 1983, comb. nov.]. However the name "Pseudomonas rhodina" corrig. has never been formally proposed before December 14, 2000 (date of publication of the amended Rule 61; see Int. J. Syst. Evol. Microbiol., 2000, 50, page 2239, column 2). Consequently, in the "List of Prokaryotic names with Standing in Nomenclature", the epithet rhodos (sic) has been maintained for Pseudomonas rhodos.

¤ Methylobacterium rhodinum corrig. (Heumann 1962) Green and Bousfield 1983, comb. nov.

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Pseudomonas rubrilineans (Lee et al. 1925) Stapp 1928, species.
Type strain: (see also StrainInfo.net) ATCC 19307= LMG 2281.
Sequence accession no. (16S rRNA gene) for the type strain: AB021421.
Synonym: "Phytomonas rubrilineans" Lee et al. 1925.
Etymology: L. adj. ruber -bra -brum, red; L. v. lineo, to reduce to a straight line, to make straight or perpendicular; N.L. part. adj. rubrilineans, making red stripes.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 LEE (H.A.), PURDY (H.A.), BARNUM (C.C.) and MARTIN (J.P.): A comparison of red-stripe disease with bacterial diseases of sugar cane and other grasses in red-stripe disease studies. Bulletin, Experiment Station of the Hawaiian Sugar Planters' Association, 1925, pp. 64-74. 2 STAPP (C.): Schizomycetes (Spaltpilze oder Bakterien). In: SORAUER (ed.), Handbuch der Pflanzenkrankheiten, 5th ed., Vol. 2, Paul Parey, Berlin, pp. 1-295.

Note: On the Approved Lists of Bacterial Names and on the Approved Lists of Bacterial Names (Amended Edition), the page numbers of the paper by Stapp are cited as pp. 758.

¤ Pseudomonas avenae Manns 1909 (Approved Lists 1980).

Note: According to Hu et al. 1991, Pseudomonas rubrilineans (Lee et al. 1925) Stapp 1928 (Approved Lists 1980) is a later heterotypic synonym of Pseudomonas avenae Manns 1909 (Approved Lists 1980).
Reference: HU (F.P.), YOUNG (J.M.) and TRIGGS (C.M.): Numerical analysis and determinative tests for nonfluorescent plant-pathogenic Pseudomonas spp. and genomic analysis and reclassification of species related to Pseudomonas avenae Manns 1909. Int. J. Syst. Bacteriol., 1991, 41, 516-525.
Original article in IJSEM Online

¤ Acidovorax avenae (Manns 1909) Willems et al. 1992, comb. nov.

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Pseudomonas rubrisubalbicans (Christopher and Edgerton 1930) Krasil'nikov 1949, species.
Type strain: (see also StrainInfo.net) ATCC 19308= CCUG 17392 = CCUG 17679 = CFBP 1202 = CIP 105019 = DSM 11543 = DSM 9440 = ICMP 5777 = JCM 21447 = LMG 2286 = NBRC 102523 = NCPPB 1027.
Sequence accession no. (16S rRNA gene) for the type strain: AF137508.
Synonym: "Phytomonas rubrisubalbicans" Christopher and Edgerton 1930.
Etymology: L. adj. ruber -bra -brum, red; L. adj. subalbicans, somewhat white, whitish; N.L. fem. adj. rubrisubalbicans, red-whitish, referring to the symptoms of mottled stripe disease.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 CHRISTOPHER (W.N.) and EDGERTON (C.W.): Bacterial stripe diseases of sugarcane in Louisiana. Journal of Agricultural Research, 1930, 41, 259-267. 2 KRASIL'NIKOV (N.A.): Guide to the bacteria and actinomycetes. Moskva-Leningrad, Izdatel'stvo Akademii Nauk SSSR, 1949, pp. 1-830.

¤ Herbaspirillum rubrisubalbicans (Christopher and Edgerton 1930) Baldani et al. 1996, comb. nov.

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Pseudomonas sabulinigri Kim et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) J64 = JCM 14963 = KCTC 22137.
Sequence accession no. (16S rRNA gene) for the type strain: EU143352.
Etymology: L. n. sabulum, sand; L. adj. niger -gra -grum, black; N.L. gen. n. sabulinigri, of black sand.
Valid publication: KIM (K.H.), ROH (S.W.), CHANG (H.W.), NAM (Y.D.), YOON (J.H.), JEON (C.O.), OH (H.M.) and BAE (J.W.): Pseudomonas sabulinigri sp. nov., isolated from black beach sand. Int. J. Syst. Evol. Microbiol., 2009, 59, 38-41.
Original article in IJSEM Online

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Pseudomonas saccharophila Doudoroff 1940, species.
Type strain: (see also StrainInfo.net) ATCC 15946= CCUG 32988 = CFBP 2433 = CIP 59.18 = DSM 654 = HAMBI 373 = IAM 14368 = JCM 21181 = LMG 2256 = NBRC 103037 = NCCB 46053 = VKM B-902.
Sequence accession no. (16S rRNA gene) for the type strain: AB021407.
Etymology: Gr. n. sakchâr, sugar; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. saccharophila, sugar-loving.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: DOUDOROFF (M.): The oxidative assimilation of sugars and related substances by Pseudomonas saccharophila with a contribution to the problem of the direct respiration of di- and polysaccharides. Enzymologia, 1940, 9, 59-72.

¤ Pelomonas saccharophila (Doudoroff 1940) Xie and Yokota 2005, comb. nov.

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Pseudomonas sagittaria Lin et al. 2013, sp. nov.
Type strain: (see also StrainInfo.net) CC-OPY-1=BCRC 80399=JCM 18195.
Sequence accession no. (16S rRNA) for the type strain: JQ277453.
Etymology: N.L. fem. adj. sagittaria (from L. n. Sagittarius the constellation Sagittarius), pertaining to the constellation Sagittarius as the novel species was isolated during the month of November, the birthday of first author.
Source: Environmental - soil.
Reference: LIN, S.-Y., HAMEED, A., LIU, Y.-C., HSU, Y.-H., LAI, W.-A., CHEN, W.-M., SHEN, F.-T. and YOUNG, C.-C. 2013. Pseudomonas sagittaria sp. nov., a siderophore-producing bacterium isolated from oil-contaminated soil. Int. J. Syst. Evol. Microbiol., 63, 2410-2417.
Original article in IJSEM Online

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Pseudomonas salegens Amoozegar et al. 2014, sp. nov.
Type strain: (see also StrainInfo.net) GBPy5=IBRC-M 10762=CECT 8338.
Sequence accession no. (16S rRNA) for the type strain: KF830707.
Etymology: L. n. sal salis, salt; L. part. adj. egens, being in need; N.L. part. adj. salegens, being in need of salt.
Source: Environmental.
Valid publication: AMOOZEGAR, M. A., SHAHINPEI, A., SEPAHY, A. A., MAKHDOUMI-KAKHKI, A., SEYEDMAHDI, S. S., SCHUMANN, P. and VENTOSA, A. 2014. Pseudomonas salegens sp. nov., a halophilic member of the genus Pseudomonas isolated from a wetland. Int. J. Syst. Evol. Microbiol., 64, 3565-3570.
Original article in IJSEM Online

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Pseudomonas salina Zhong et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) XCD-X85=CGMCC 1.12482=JCM 19469.
Sequence accession no. (16S rRNA) for the type strain: KC762324.
Etymology: L. fem. adj. salina, salty.
Source: Environmental.
Valid publication: ZHONG, Z. P., LIU, Y., HOU, T. T., LIU, H. C., ZHOU, Y. G., WANG, F. and LIU, Z. P. 2015. Pseudomonas salina sp. nov., isolated from a salt lake. Int. J. Syst. Evol. Microbiol., 65, 2846-2851.
Original article in IJSEM Online

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Pseudomonas salomonii Gardan et al. 2002, sp. nov.
Type strain: (see also StrainInfo.net) CFBP 2022 = ICMP 14252 = NCPPB 4277.
Sequence accession no. (16S rRNA gene) for the type strain: AY091528.
Etymology: N.L. gen. masc. n. salomonii, of Salomon, referring to Max Salomon, the agricultural technical advisor who firstobserved the symptoms of the new garlic disease "Café au lait".
Valid publication: GARDAN (L.), BELLA (P.), MEYER (J.M.), CHRISTEN (R.), ROTT (P.), ACHOUAK (W.) and SAMSON (R.): Pseudomonas salomonii sp. nov., pathogenic on garlic, and Pseudomonas palleroniana sp. nov., isolated from rice. Int. J. Syst. Evol. Microbiol., 2002, 52, 2065-2074.
Original article in IJSEM Online

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Pseudomonas saponiphila Lang et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) CIP 109856 = DSM 9751.
Sequence accession no. (16S rRNA gene) for the type strain: FM208264.
Etymology: L. n. sapo -onis, soap; N.L. fem. adj. phila (from Gr. fem. adj. philê), friend, loving; N.L. fem. adj. saponiphila, soap (detergent)- loving.
Valid publication: Validation List no. 145. Int. J. Syst. Evol. Microbiol., 2012, 62, 1017-1019.
Effective publication: LANG (E.), BURGHARTZ (M.), SPRING (S.), SWIDERSKI (J.) and SPRÖER (C.): Pseudomonas benzenivorans sp. nov. and Pseudomonas saponiphila sp. nov., represented by xenobiotics degrading type strains. Curr. Microbiol., 2010, 60, 85-91.
Validation List no. 145 in IJSEM Online - Effective publication Online

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Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992, comb. nov.
Type strain: (see also StrainInfo.net) ATCC 13522= CFBP 1670 = CIP 103721 = ICMP 4352 = LMG 2209 = NCPPB 639.
Sequence accession no. (16S rRNA gene) for the type strain: AB021402.
Basonym: ¤ Pseudomonas syringae subsp. savastanoi (ex Smith 1908) Janse 1982.
Other synonym: "Pseudomonas savastanoi" (Smith 1908) Stevens 1913.
Etymology: N.L. gen. masc. n.savastanoi, of Savastano, the first worker who studied olive knot disease.
Valid publication: GARDAN (L.), BOLLET (C.), ABU GHORRAH (M.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovar strains of Pseudomonas syringae subsp. savastanoi Janse (1982) and proposal of Pseudomonas savastanoi sp. nov. Int. J. Syst. Bacteriol., 1992, 42, 606-612.
Original article in IJSEM Online

Note: Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992, ¤ Pseudomonas amygdali Psallidas and Panagopoulos 1975 (Approved Lists 1980), ¤ Pseudomonas ficuserectae Goto 1983, and ¤ Pseudomonas meliae Ogimi 1981, and are members of a single genomospecies. Thus, these species should be considered synonymous and the correct name for this species should be Pseudomonas amygdali. Unfortunately, Pseudomonas amygdali, as delineated by DNA-DNA hybridization, could not be differentiated from other genomospecies by phenotypic tests. So the authors do not formally propose nomenclatural changes.
Reference: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Sutic and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

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Pseudomonas segetis Park et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) FR1439 = CIP 108523 = IMSNU 14100 = KCTC 12331.
Sequence accession no. (16S rRNA gene) for the type strain: AY770691.
Etymology: L. n. seges -etis, soil; L. gen. n. segetis, of the soil.
Valid publication: PARK (Y.D.), YI (H.), BAIK (K.S.), SEONG (C.N.), BAE (K.S.), MOON (E.Y.) and CHUN (J.): Pseudomonas segetis sp. nov., isolated from soil. Int. J. Syst. Evol. Microbiol., 2006, 56, 2593-2595.
Original article in IJSEM Online

Note: In the paper by Park et al. 2006, the type strain IMSNU 14100 is erroneously cited as IMSNU 14101.

Pseudomonas segetis Park et al. 2006 emend. Hameed et al. 2014.
Reference: HAMEED, A., SHAHINA, M., LIN, S.-Y., LIU, Y.-C. and YOUNG, C.-C. 2014. Pseudomonas hussainii sp. nov., isolated from droppings of a seashore bird, and emended descriptions of Pseudomonas pohangensis, Pseudomonas benzenivorans and Pseudomonas segetis. Int. J. Syst. Evol. Microbiol., 64, 2330-2337.
Original article.

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Pseudomonas seleniipraecipitans corrig. Hunter and Manter 2011, sp. nov.
Type strain: (see also StrainInfo.net) CA5 = LMG 25475 = NRRL B-51283.
Sequence accession no. (16S rRNA gene) for the type strain: FJ422810.
Etymology: N.L. n. selenium selenium; L. part. adj. praecipitans precipitating; N.L. part. adj. seleniipraecipitans selenium precipitating; referring to the organism's ability to remove the selenium oxyanion selenite from aqueous solution.
Valid publication: Validation List no. 140. Int. J. Syst. Evol. Microbiol., 2011, 61, 1499-1501.
Effective publication: HUNTER (W.J.) and MANTER (D.K.): Pseudomonas seleniipraecipitatus sp. nov.: a selenite reducing γ-proteobacteria isolated from soil. Curr. Microbiol., 2011, 62, 565-569.
Validation List no. 140 in IJSEM Online - Effective publication Online

Note: The original spelling of the specific epithet, seleniipraecipitatus (sic), has been corrected on validation according to Rule 61.
Reference: ASSOCIATE EDITOR, IJSEM: Validation List no. 140 (footnote ##). Int. J. Syst. Evol. Microbiol., 2011, 61, 1499-1501.
Validation List no. 140 in IJSEM Online

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Pseudomonas simiae Vela et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) OLi = CCUG 50988 = CECT 7078.
Sequence accession no. (16S rRNA gene) for the type strain: AJ936933.
Etymology: L. gen. n. simiae, of an ape, of a monkey.
Valid publication: VELA (A.I.), GUTIÉRREZ (M.C.), FALSEN (E.), ROLLÁN (E.), SIMARRO (I.), GARCÍA (P.), DOMÍNGUEZ (L.), VENTOSA (A.) and FERNÁNDEZ-GARAYZÁBAL (J.F.): Pseudomonas simiae sp. nov., isolated from clinical specimens from monkeys (Callithrix geoffroyi). Int. J. Syst. Evol. Microbiol., 2006, 56, 2671-2676.
Original article in IJSEM Online

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Pseudomonas solanacearum (Smith 1896) Smith 1914, species.
Type strain: (see also StrainInfo.net) A. Kelman 60-1 = ATCC 11696= CCUG 14272 = CFBP 2047 = CIP 104762 = DSM 9544 = ICMP 5712 = JCM 10489 = LMG 2299 = NCAIM B.01459 = NCPPB 325 = NRRL B-3212.
Sequence accession no. (16S rRNA gene) for the type strain: EF016361.
Synonym: "Bacillus solanacearum" Smith 1896.
Etymology: N.L. fem. pl. n. Solanaceae, the nightshade family; N.L. fem. pl. gen. n. solanacearum, of the Solanaceae.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SMITH (E.F.): A bacterial disease of the tomato, eggplant and Irish potato (Bacillus solanacearum n. sp.). Bulletin, Division of Vegetable Physiology and Pathology, United States Department of Agriculture, 1896, 12, 1-28. 2 SMITH (E.F.): Bacteria in relation to plant diseases. Carnegie Institute, Washington, 1914, 3, 1-309.

¤ Burkholderia solanacearum (Smith 1896) Yabuuchi et al. 1993, comb. nov.

¤ Ralstonia solanacearum (Smith 1896) Yabuuchi et al. 1996, comb. nov.

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Pseudomonas soli Pascual et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) F-279,208=DSM 28043=LMG 27941.
Sequence accession no. (16S rRNA) for the type strain: HF930598.
Whole-genome sequence accession no. for the type strain: -.
Etymology: -.
Source: Environmental - soil.
Valid publication: Validation List no. 162. List of new names and new combinations previously effectively, but not validly, published. Int. J Syst. Evol. Microbiol., 65 (2015), 741-744.
Validation List Online
Effective publication: PASCUAL, J., GARCÍA-LÓPEZ, M., CARMONA, C., DA S. SOUSA, T., DE PEDRO, N., CAUTAIN, B., MARTÍN, J., VICENTE, F., REYES, F., BILLS, G. F. and GENILLOUD, O. 2014. Pseudomonas soli sp. nov., a novel producer of xantholysin congeners. Syst. Appl. Microbiol., 37, 412-416.
Effective publication Online

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Pseudomonas spinosa Leifson 1962, species.
Type strain: (see also StrainInfo.net) 83 = ATCC 14606= DSM 15801 = JCM 21400.
Sequence accession no. (16S rRNA gene) for the type strain: AB077038.
Etymology: L. fem. adj. spinosa, thorny, spiny.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: LEIFSON (E.): Pseudomonas spinosa n. sp. International Bulletin of Bacteriological Nomenclature and Taxonomy, 1962, 12, 89-92.

¤ Malikia spinosa (Leifson 1962) Spring et al. 2005, comb. nov.

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Pseudomonas stanieri Baumann et al. 1983, sp. nov.
Type strain: (see also StrainInfo.net) 146 = ATCC 27130= CCUG 16021 = CIP 106697 = DSM 7027 = LMG 6847.
Sequence accession no. (16S rRNA gene) for the type strain: AB021367.
Etymology: N.L. gen. masc. n. stanieri, of Stanier, named after R. Y. Stanier, a Canadian microbiologist who made extensive contributions to general microbiology and, in particular, to Pseudomonas.
Valid publication: BAUMANN (P.), BOWDITCH (R.D.), BAUMANN (L.) and BEAMAN (B.): Taxonomy of marine Pseudomonas species: P. stanieri sp. nov.; P. perfectomarina sp. nov., nom. rev.; P. nautica; and P. doudoroffii. Int. J. Syst. Bacteriol., 1983, 33, 857-865.
Original article in IJSEM Online

¤ Marinobacterium stanieri (Baumann et al. 1983) Satomi et al. 2002, comb. nov.

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Pseudomonas straminea corrig. Iizuka and Komagata 1963, species.
Type strain: (see also StrainInfo.net) ATCC 33636= CCUG 12539 = CIP 106745 = IAM 1598 = IFO (now NBRC) 16665 = JCM 2783 = NRIC 0164.
Sequence accession no. (16S rRNA gene) for the type strain: D84023.
Etymology: L. fem. adj. straminea, made of straw.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: IIZUKA (H.) and KOMAGATA (K.): New species of Pseudomonas belonged to fluorescent group (Studies on the microorganisms of cereal grains. Part V). Journal of the Agricultural Chemical Society of Japan, 1963, 37, 137-141.

Note: The original spelling of the specific epithet, straminae (sic), has been corrected by Uchino et al. 2000.
Reference: UCHINO (M.), KOSAKO (Y.), UCHIMURA (T.) and KOMAGATA (K.): Emendation of Pseudomonas straminea Iizuka and Komagata 1963. Int. J. Syst. Evol. Microbiol., 2000, 50, 1513-1519.
Original article in IJSEM Online

Pseudomonas straminea corrig. Iizuka and Komagata 1963 (Approved Lists 1980) emend. Uchino et al. 2000.
Type strain: (see also StrainInfo.net) ATCC 33636= CIP 106745 = IAM 1598 = JCM 2783 = NRIC 0164.
Sequence accession no. (16S rRNA gene) for the type strain: D84023.
Etymology: L. fem. adj. straminea, made of straw.
Reference: UCHINO (M.), KOSAKO (Y.), UCHIMURA (T.) and KOMAGATA (K.): Emendation of Pseudomonas straminea Iizuka and Komagata 1963. Int. J. Syst. Evol. Microbiol., 2000, 50, 1513-1519.
Original article in IJSEM Online

Note: Pseudomonas straminea corrig. Iizuka and Komagata 1963 (Approved Lists 1980) emend. Uchino et al. 2000 includes the strains of "Pseudomonas ochracea" (Zimmermann 1890) Chester 1901.

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Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946, species.
Type strain: (see also StrainInfo.net) ATCC 17588= CCUG 11256 = CFBP 2443 = CIP 103022 = DSM 5190 = IFO (now NBRC) 14165 = JCM 5965 = LMG 11199 = NCCB 76042 = VKM B-975.
Sequence accession no. (16S rRNA gene) for the type strain: AF094748, CP002881 (complete genome).
Synonyms: "Bacillus denitrificans II" Burri and Stutzer 1895, "Bacterium stutzeri" Lehmann and Neumann 1896, "Bacillus nitrogenes" Migula 1900, "Bacillus stutzeri" Chester 1901, "Achromobacter sewerinii" Bergey et al. 1923, "Achromobacter stutzeri" Bergey et al. 1930, "Pseudomonas stanieri" Mandel 1966, not "Pseudomonas stutzeri" Migula 1900.
Etymology: N.L. gen. masc. n. stutzeri, of Stutzer.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 LEHMANN (K.B.) and NEUMANN (R.): Atlas und Grundriss der Bakteriologie und Lehrbuch der speziellen bakteriologischen Diagnostik, 1st ed., J.F. Lehmann, München, 1896. 2 SIJDERIUS (R.): Heterotrophe bacterien, die thiosulfaat oxydeeren. Thesis, University Amsterdam, 1946, pp. 1-146.

Notes:
1 According to Döhler et al. 1987, Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas perfectomarina (ex ZoBell and Upham 1944) Baumann et al. 1983.
Reference: DÖHLER (K.), HUSS (V.A.R.) and ZUMFT (W.G.): Transfer of Pseudomonas perfectomarina Baumann, Bowditch, Baumann, and Beaman 1983 to Pseudomonas stutzeri (Lehmann and Neumann 1896) Sidjerius 1946. Int. J. Syst. Bacteriol., 1987, 37, 1-3.
Original article in IJSEM Online
2 According to Cladera et al. 2006, Pseudomonas stutzeri (Lehmann and Neumann 1896) Sijderius 1946 (Approved Lists 1980) is an earlier heterotypic synonym of ¤ Pseudomonas chloritidismutans Wolterink et al. 2002.
References: List of Changes in Taxonomic Opinion no. 4. Int. J. Syst. Evol. Microbiol., 2006, 56, 1463. [CLADERA (A.M.), GARCÍA-VALDÉS (E.) and LALUCAT (J.): Genotype versus phenotype in the circumscription of bacterial species: the case of Pseudomonas stutzeri and Pseudomonas chloritidismutans. Arch. Microbiol., 2006, 184, 353-361.]
List of Changes in Taxonomic Opinion no. 4 in IJSEM Online - Effective publication Online

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Pseudomonas synxantha (Ehrenberg 1840) Holland 1920, species.
Type strain: (see also StrainInfo.net) ATCC 9890= CFBP 5591 = CIP 59.22 = IFO (now NBRC) 3913 = JCM 11599 = LMG 2190 = NCTC 10696 = NRRL B-780.
Sequence accession no. (16S rRNA gene) for the type strain: D84025.
Synonym: "Vibrio synxanthus" Ehrenberg 1840.
Etymology: Gr. pref. sun, with, along with, together; Gr. adj. xanthos, yellow; N.L. fem. adj. synxantha, with yellow.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 EHRENBERG (C.G.): Charakteristik von 274 neuen Arten von Infusorien. Bericht Bekannt Verhandlungen der Königlisch-Preussichen Akademie der Wissenschaftlichen, Berlin, 1840, 197-219. 2 HOLLAND (D.F.): V. Generic index of the commoner forms of bacteria (p. 215). In:. WINSLOW (C.E.A.), BROADHURST (J.), BUCHANAN (R.E.), KRUMWIEDE (C.) Jr., ROGERS (L.A.) and SMITH (G.H.): The families and genera of the bacteria. Journal of Bacteriology, 1920, 5, 191-229.

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Pseudomonas syringae van Hall 1902, species.
Type strain: (see also StrainInfo.net) ATCC 19310= CCUG 14279 = CFBP 1392 = CIP 106698 = ICMP 3023 = LMG 1247 = NCAIM B.01398 = NCPPB 281 = NRRL B-1631.
Sequence accession no. (16S rRNA gene) for the type strain: DQ318866.
Etymology: N.L. fem. n. Syringa, generic name of lilac; N.L. fem. gen. n. syringae, of the lilac.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: VAN HALL (C.J.J.): Bijdragen tot de kennis der Bakterieele Plantenziekten. Inaugural Dissertation Amsterdam, 1902, pp. 198.

Notes:
1 Pseudomonas syringae van Hall 1902 (Approved Lists 1980) as it is presently circumscribed is represented by seven genomospecies which may be considered to represent species.
Reference: YOUNG (J.M.), SADDLER (G.S.), TAKIKAWA (Y.), DE BOER (S.H.), VAUTERIN (L.), GARDAN (L.), GVOZDYAK (R.I.) and STEAD (D.E.): Names of plant pathogenic bacteria 1864-1995. Review of plant pathology, 1996, 75, 721-763.
2 Subsequently, this species has been divided into subspecies (see: below).

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Pseudomonas syringae subsp. savastanoi (ex Smith 1908) Janse 1982, subsp. nov., nom. rev.
Type strain: (see also StrainInfo.net) ATCC 13522= CFBP 1670 = CIP 103721 = ICMP 4352 = LMG 2209 = NCPPB 639.
Sequence accession no. (16S rRNA gene) for the type strain: AB021402.
Synonym: "Pseudomonas savastanoi" (Smith 1908) Stevens 1913.
Etymology: N.L. gen. masc. n.savastanoi, of Savastano, the first worker who studied olive knot disease.
Valid publication: JANSE (J.D.): Pseudomonas syringae subsp. savastanoi (ex Smith) subsp. nov., nom. rev., the bacterium causing excrescences on Oleaceae and Nerium oleander L. Int. J. Syst. Bacteriol., 1982, 32, 166-169.
Original article in IJSEM Online

¤ Pseudomonas savastanoi (Janse 1982) Gardan et al. 1992, comb. nov.

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Pseudomonas syringae subsp. syringae (van Hall 1902) Janse 1982, subsp. nov.
Type strain: (see also StrainInfo.net) ATCC 19310= CCUG 14279 = CFBP 1392 = CIP 106698 = ICMP 3023 = LMG 1247 = NCAIM B.01398 = NCPPB 281.
Sequence accession no. (16S rRNA gene) for the type strain: DQ318866.
Etymology: N.L. fem. n. Syringa, generic name of lilac; N.L. fem. gen. n. syringae, of the lilac.
Valid publication: HOWEY (R.T.), LOCK (C.M.) and MOORE (L.V.H.): Subspecies names automatically created by Rule 46. Int. J. Syst. Bacteriol., 1990, 40, 317-319.
Original article in IJSEM Online

Notes:
1 The original authorship, van Hall 1902, was corrected according to the new Rule 40d.
Reference: GARRITY (G.M.), LABEDA (D.P.) and OREN (A.): Judicial Commission of the International Committee on Systematics of Prokaryotes. XIIth International (IUMS) Congress of Bacteriology and Applied Microbiology. Minutes of the meetings, 3, 4 and 6 August 2008, Istanbul, Turkey. Int. J. Syst. Evol. Microbiol., 2011, 61, 2775-2780.
Original article in IJSEM Online
2 The subspecies name Pseudomonas syringae subsp. syringae (van Hall 1902) Janse 1982 is automatically created by the valid publication of Pseudomonas syringae subsp. savastanoi (ex Smith 1908) Janse 1982 [Rule 40d (formerly Rule 46)].
3 If an author accepts the elevation of Pseudomonas syringae subsp. savastanoi (ex Smith 1908) Janse 1982 to full species status, then Pseudomonas syringae subsp. syringae (van Hall 1902) Janse 1982 should thereafter be designated simply as Pseudomonas syringae van Hall 1902 (Approved Lists 1980).

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Pseudomonas syzygii Roberts et al. 1990, sp. nov.
Type strain: (see also StrainInfo.net) R001 = ATCC 49543= CCUG 32781 = DSM 7385 = ICMP 10915 = LMG 10661 = NCPPB 3446.
Sequence accession no. (16S rRNA gene) for the type strain: AB021403.
Etymology: N.L. n. Syzygium, generic name of the clove tree; L. gen. n. syzygii, of the genus Syzygium.
Valid publication: Validation List no. 34. Int. J. Syst. Bacteriol., 1990, 40, 320-321.
Effective publication: ROBERTS (S.J.), EDEN-GREEN (S.J.), JONES (P.) and AMBLER (D.J.): Pseudomonas syzygii sp. nov., the cause of Sumatra disease of cloves. Syst. Appl. Microbiol., 1990, 13, 34-43.
Validation List no. 34 in IJSEM Online - Effective publication Online

¤ Ralstonia syzygii (Roberts et al. 1990) Vaneechoutte et al. 2004, comb. nov.

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Pseudomonas taeanensis Lee et al. 2010, sp. nov.
Type strain: (see also StrainInfo.net) MS-3 = JCM 16046 = KACC 14032 = KCTC 22612 = NBRL 105641.
Sequence accession no. (16S rRNA gene) for the type strain: FJ424813.
Etymology: N.L. fem. adj. taeanensis, of or pertaining to Taean, Korea, where the type strain was isolated.
Valid publication: LEE (D.H.), MOON (S.R.), PARK (Y.H.), KIM (J.H.), KIM (H.), PARALES (R.E.) and KAHNG (H.Y.): Pseudomonas taeanensis sp. nov., isolated from a crude oil-contaminated seashore. Int. J. Syst. Evol. Microbiol., 2010, 60, 2719-2723.
Original article in IJSEM Online

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Pseudomonas taeniospiralis Lalucat et al. 1982, sp. nov.
Type strain: (see also StrainInfo.net) 2K1 = ATCC 49743= CIP 106727 = CCUG 15921 = DSM 2082 = JCM 21411 = LMG 7170 = NBRC 102512.
Sequence accession no. (16S rRNA gene) for the type strain: AF078768.
Etymology: Gr. n. tainia (Latin transliteration taenia), band, ribbon; N.L. fem. adj. spiralis (from L. n. spira, coil; and L. fem. suff. -alis, suffix denoting pertaining to), coiled; N.L. fem. adj. taeniospiralis, ribbon coiled, after Caedibacter taeniospiralis, an organism with which it shares characteristics.
Valid publication: LALUCAT (J.), PARÉS (R.) and SCHLEGEL (H.G.): Pseudomonas taeniospiralis sp. nov., an R-body-containing hydrogen bacterium. Int. J. Syst. Bacteriol., 1982, 32, 332-338.
Original article in IJSEM Online

¤ Hydrogenophaga taeniospiralis (Lalucat et al. 1982) Willems et al. 1989, comb. nov.

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Pseudomonas taetrolens Haynes 1957, species.
Type strain: (see also StrainInfo.net) ATCC 4683= CCUG 560 = CFBP 5592 = CIP 103299 = JCM 20238 = LMG 2336 = NCTC 10697 = NRRL B-14 = NRRL B-731 = VKM B-904.
Sequence accession no. (16S rRNA gene) for the type strain: D84027.
Etymology: L. adj. taeter -tra -trum, offensive, foul; L. part. adj. olens, having an odor; N.L. part. adj. taetrolens, foul smelling.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: HAYNES (W.C.): Genus I. Pseudomonas Migula 1894. In: R.S. BREED, E.G.D. MURRAY and N.R. SMITH (editors): Bergey's Manual of Determinative Bacteriology, 7th edition, The Williams & Wilkins Co, Baltimore, 1957, pp. 89-152.

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Pseudomonas taiwanensis Wang et al. 2010, sp. nov.
Type strain: (see also StrainInfo.net) CMS = BCRC 17751 = DSM 21245.
Sequence accession no. (16S rRNA gene) for the type strain: EU103629.
Etymology: N.L. fem. adj. taiwanensis, of or pertaining to Taiwan, where the type strain was isolated.
Valid publication: WANG (L.T.), TAI (C.J.), WU (Y.C.), CHEN (Y.B.), LEE (F.L.) and WANG (S.L.): Pseudomonas taiwanensis sp. nov., isolated from soil. Int. J. Syst. Evol. Microbiol., 2010, 60, 2094-2098.
Original article in IJSEM Online

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Pseudomonas testosteroni Marcus and Talalay 1956, species.
Type strain: (see also StrainInfo.net) KS 0043 = ATCC 11996= CCUG 1426 = CFBP 2436 = CIP 59.24 = DSM 50244 = HAMBI 403 = IAM 12419 = IFO (now NBRC) 14951 = JCM 5832 = JCM 6221 = LMG 1786 = LMG 1800 = NCAIM B.01655 = NCCB 86053 = NCIB (now NCIMB) 8955 = NCTC 10698 = NRRL B-2611 = VKM B-1241.
Sequence accession no. (16S rRNA gene) for the type strain: M11224.
Etymology: N.L. gen. n. testosteroni, of testosterone, a chemical compound.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: MARCUS (P.I.) and TALALAY (P.): Induction and purification of α- and β-hydroxysteroid dehydrogenases. Journal of Biological Chemistry, 1956, 218, 661-674.

¤ Comamonas testosteroni (Marcus and Talalay 1956) Tamaoka et al. 1987, comb. nov.

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Pseudomonas thermotolerans Manaia and Moore 2002, sp. nov.
Type strain: (see also StrainInfo.net) CM3 = DSM 14292 = LMG 21284.
Sequence accession no. (16S rRNA gene) for the type strain: AJ311980.
Etymology: Gr. n. thermê, heat; L. part. adj. tolerans, tolerating; N.L. part. adj. thermotolerans, able to tolerate high temperatures.
Valid publication: MANAIA (C.M.) and MOORE (E.R.B.): Pseudomonas thermotolerans sp. nov., a thermotolerant species of the genus Pseudomonas sensu stricto. Int. J. Syst. Evol. Microbiol., 2002, 52, 2203-2209.
Original article in IJSEM Online

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Pseudomonas thivervalensis Achouak et al. 2000, sp. nov.
Type strain: (see also StrainInfo.net) SBK26 = CFBP 11261 = DSM 13194 = JCM 11941.
Sequence accession no. (16S rRNA gene) for the type strain: AF100323.
Etymology: N.L. fem. adj. thivervalensis, of or belonging to Thiverval, France, the place where most of the strains have been isolated from the rhizoplane of Arabidopsis ².
Valid publication: ACHOUAK (W.), SUTRA (L.), HEULIN (T.), MEYER (J.M.), FROMIN (N.), DEGRAEVE (S.), CHRISTEN (R.) and GARDAN (L.): Pseudomonas brassicacearum sp. nov. and Pseudomonas thivervalensis sp. nov., two root-associated bacteria isolated from Brassica napus and Arabidopsis thaliana. Int. J. Syst. Evol. Microbiol., 2000, 50, 9-18.
Original article in IJSEM Online

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Pseudomonas tolaasii Paine 1919, species.
Type strain: (see also StrainInfo.net) ATCC 33618= CCUG 23369 = CCUG 32782 = CFBP 2068 = CIP 106735 = ICMP 12833 = JCM 21583 = LMG 2342 = NBRC 103163 = NCPPB 2192.
Sequence accession no. (16S rRNA gene) for the type strain: AF255336.
Etymology: N.L. gen. masc. n. tolaasii, of Tolaas.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: PAINE (S.G.): Studies in bacteriosis. II. A brown blotch disease of cultivated mushrooms. Annals of Applied Biology, 1919, 5, 206-219.

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Pseudomonas toyotomiensis Hirota et al. 2011, sp. nov.
Type strain: (see also StrainInfo.net) HT-3 = JCM 15604 = NCIMB 14511.
Sequence accession no. (16S rRNA gene) for the type strain: AB453701.
Etymology: N.L. fem. adj. toyotomiensis, of or pertaining to Toyotomi, where the type strain was isolated.
Valid publication: HIROTA (K.), YAMAHIRA (K.), NAKAJIMA (K.), NODASAKA (Y.), OKUYAMA (H.) and YUMOTO (I.): Pseudomonas toyotomiensis sp. nov., a psychrotolerant facultative alkaliphile that utilizes hydrocarbons. Int. J. Syst. Evol. Microbiol., 2011, 61, 1842-1848.
Original article in IJSEM Online

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Pseudomonas tremae Gardan et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) CFBP 3229 = CIP 106417 = ICMP 9151 = NCPPB 3465.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492826.
Etymology: N.L. gen. fem. n. tremae, of Trema; generic name of the host plant, Trema orientalis BL.
Valid publication: GARDAN (L.), SHAFIK (H.), BELOUIN (S.), BROCH (R.), GRIMONT (F.) and GRIMONT (P.A.D.): DNA relatedness among the pathovars of Pseudomonas syringae and description of Pseudomonas tremae sp. nov. and Pseudomonas cannabina sp. nov. (ex Sutic and Dowson 1959). Int. J. Syst. Bacteriol., 1999, 49, 469-478.
Original article in IJSEM Online

Note: Pseudomonas tremae Gardan et al. 1999 includes Pseudomonas syringae pv. Tremae.

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Pseudomonas trivialis Behrendt et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) P 513/19 = DSM 14937 = LMG 21464.
Sequence accession no. (16S rRNA gene) for the type strain: AJ492831.
Etymology: L. fem. adj. trivialis, trivial.
Valid publication: BEHRENDT (U.), ULRICH (A.) and SCHUMANN (P.): Fluorescent pseudomonads associated with the phyllosphere of grasses; Pseudomonas trivialis sp. nov., Pseudomonas poae sp. nov. and Pseudomonas congelans sp. nov. Int. J. Syst. Evol. Microbiol., 2003, 53, 1461-1469.
Original article in IJSEM Online

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Pseudomonas tuomuerensis Xin et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) 78-123 = CGMCC 1.1365 = JCM 14085.
Sequence accession no. (16S rRNA gene) for the type strain: DQ868767.
Etymology: N.L. fem. adj. tuomuerensis, pertaining to the region of Tuomuer Peak of Tianshan Mountain, where the type strain was isolated.
Valid publication: XIN (Y.H.), ZHANG (D.C.), LIU (H.C.), ZHOU (H.L.) and ZHOU (Y.G.): Pseudomonas tuomuerensis sp. nov., isolated from a bird's nest. Int. J. Syst. Evol. Microbiol., 2009, 59, 139-143.
Original article in IJSEM Online

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Pseudomonas turukhanskensis Korshunova et al. 2016, sp. nov.
Type strain: (see also StrainInfo.net) IB1.1=VKM B-2935=CECT 9091.
Sequence accession no. (16S rRNA) for the type strain: KP306892.
Whole-genome sequence accession no. for the type strain: .
Etymology: N.L. fem. adj. turukhanskensis, pertaining to Turukhansk District, the name of the administrative unit of Krasnoyarsk Krai (East Siberia, Russia) where the type strain was isolated.
Source: Environmental - soil.
Valid publication: KORSHUNOVA, T. Y., RAMIREZ-BAHENA, M. H., CHETVERIKOV, S. P., IGUAL, J. M., PEIX, A. and LOGINOV, O. 2016. Pseudomonas turukhanskensis sp. nov., isolated from oil-contaminated soils. Int. J. Syst. Evol. Microbiol., 66, 4657-4664.
Original article in IJSEM Online

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Pseudomonas umsongensis Kwon et al. 2003, sp. nov.
Type strain: (see also StrainInfo.net) Ps 3-10 = KACC 10847 = LMG 21317.
Sequence accession no. (16S rRNA gene) for the type strain: AF468450.
Etymology: N.L. fem. adj. umsongensis, referring to Umsong Region in Korea, where the bacteria were first found.
Valid publication: KWON (S.W.), KIM (J.S.), PARK (I.C.), YOON (S.H.), PARK (D.H.), LIM (C.K.) and GO (S.J.): Pseudomonas koreensis sp. nov., Pseudomonas umsongensis sp. nov. and Pseudomonas jinjuensis sp. nov., novel species from farm soils in Korea. Int. J. Syst. Evol. Microbiol., 2003, 53, 21-27.
Original article in IJSEM Online

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Pseudomonas vancouverensis Mohn et al. 1999, sp. nov.
Type strain: (see also StrainInfo.net) DhA-51 = ATCC 700688= CCUG 49675 = CIP 106707.
Sequence accession no. (16S rRNA gene) for the type strain: AJ011507.
Etymology: N.L. fem. adj. vancouverensis, pertaining to the city of Vancouver, Canada.
Valid publication: Validation List no. 70. Int. J. Syst. Bacteriol., 1999, 49, 935-936.
Effective publication: MOHN (W.W.), WILSON (A.E.), BICHO (P.) and MOORE (E.R.B.): Physiological and phylogenetic diversity of bacteria growing on resin acids. Syst. Appl. Microbiol., 1999, 22, 68-78.
Validation List no. 70 in IJSEM Online - Effective publication Online

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Pseudomonas veronii Elomari et al. 1996, sp. nov.
Type strain: (see also StrainInfo.net) ATCC 700272= ATCC 700474= CCUG 43519 = CIP 104663 = CFML 92-134 = DSM 11331 = JCM 11942 = LMG 17761.
Sequence accession no. (16S rRNA gene) for the type strain: AF064460.
Etymology: N.L. masc. gen. n. veronii, of Véron, in honor of Prof. M. M. Véron, an eminent French microbiologist, for his contribution to taxonomy and medical microbiology.
Valid publication: ELOMARI (M.), COROLER (L.), HOSTE (B.), GILLIS (M.), IZARD (D.) and LECLERC (H.): DNA relatedness among Pseudomonas strains isolated from natural mineral waters and proposal of Pseudomonas veronii sp. nov. Int. J. Syst. Bacteriol., 1996, 46, 1138-1144.
Original article in IJSEM Online

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Pseudomonas vesicularis (Büsing et al. 1953) Galarneault and Leifson 1964, species.
Type strain: (see also StrainInfo.net) ATCC 11426= CCM 3398 = BCRC (formerly CCRC) 11012 = CCUG 2032 = CCUG 28332 = CCUG 28333 = CFBP 3032 = CIP 101035 = DSM 7226 = DSM 7233 = IAM 12105 = IFO (now NBRC) 12165 = JCM 1477 = LMG 2350 = NCCB 76049 = NCIMB 1945 = NCTC 10900 = VKM B-974.
Sequence accession no. (16S rRNA gene) for the type strain: AJ227780.
Synonyms: "Corynebacterium vesiculare" Büsing et al. 1953, "Pseudomonas vesiculare" (sic) (Büsing et al. 1953) Galarneault and Leifson 1964.
Etymology: L. n. vesicula, a little blister, vesicle; L. fem. suff. -aris, suffix denoting pertaining to; N.L. fem. adj. vesicularis, pertaining to a vesicle.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 BUSING (K.H.), DOLL (W.) and FREYTAG (K.): Die Bakterienflora der medizinischen Blutegel. Archiv fur Mikrobiologie, 1953, 19, 52-86. 2 GALARNEAULT (T.P.) and LEIFSON (E.): Pseudomonas vesiculare (Büsing et al.) nov. comb. International Bulletin of Bacteriological Nomenclature and Taxonomy, 1964, 14, 165-168.

¤ Brevundimonas vesicularis (Büsing et al. 1953) Segers et al. 1994, comb. nov.

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Pseudomonas viridiflava (Burkholder 1930) Dowson 1939, species.
Type strain: (see also StrainInfo.net) ATCC 13223= CFBP 2107 = CIP 106699 = DSM 11124 = ICMP 2848 = LMG 2352 = NCPPB 635 = NRRL B-895.
Sequence accession no. (16S rRNA gene) for the type strain: AY180972.
Synonym: "Phytomonas viridiflava" Burkholder 1930.
Etymology: L. adj. viridis, green; L. adj. flavus, golden yellow, reddish yellow; N.L. fem. adj. viridiflava, greenish yellow.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 BURKHOLDER (W.H.): The bacterial diseases of the bean. Memoirs, Cornell University Agricultural Experiment Station, 1930, 127, 1-88. 2 DOWSON (W.J.): On the systematic position and generic names of the Gram negative bacterial plant pathogens. Zentralblatt fur Bakteriologie, Parasitenkunde, Infektionskrankheiten und Hygiene. Abteilung II, 1939, 100, 177-193.

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Pseudomonas vranovensis Tvrzová et al. 2006, sp. nov.
Type strain: (see also StrainInfo.net) 2B2 = CCM 7279 = DSM 16006.
Sequence accession no. (16S rRNA gene) for the type strain: AY970951.
Etymology: N.L. fem. adj. vranovensis, pertaining to Vranov, the name of a village in South Moravia, the place of soil origin, the source of isolation of strain 2B2.
Valid publication: TVRZOVÁ (L.), SCHUMANN (P.), SPRÖER (C.), SEDLÁČEK (I.), PÁČOVÁ (Z.), ŠEDO (O.), ZDRÁHAL (Z.), STEFFEN (M.) and LANG (E.): Pseudomonas moraviensis sp. nov. and Pseudomonas vranovensis sp. nov., soil bacteria isolated on nitroaromatic compounds, and emended description of Pseudomonas asplenii. Int. J. Syst. Evol. Microbiol., 2006, 56, 2657-2663.
Original article in IJSEM Online

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Pseudomonas weihenstephanensis Von Neubeck et al. 2016, sp. nov.
Type strain: (see also StrainInfo.net) WS4993=DSM 29166=LMG 28437.
Sequence accession no. (16S rRNA) for the type strain: KP738720.
Whole-genome sequence accession no. for the type strain: -.
Etymology: N.L. fem. adj. weihenstephanensis, pertaining to Freising/Weihenstephan in southern Germany, where the type strain was isolated.
Source: Animal.
Valid publication: VON NEUBECK, M., HUPTAS, C., GLUCK, C., KREWINKEL, M., STOECKEL, M., STRESSLER, T., FISCHER, L., HINRICHS, J., SCHERER, S. and WENNING, M. 2016. Pseudomonas helleri sp. nov. and Pseudomonas weihenstephanensis sp. nov., isolated from raw cow's milk. Int. J. Syst. Evol. Microbiol., 66, 1163-1173.
Original article in IJSEM Online

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Pseudomonas woodsii (Smith 1911) Stevens 1925, species.
Type strain: (see also StrainInfo.net) ATCC 19311= LMG 2362.
Sequence accession no. (16S rRNA gene) for the type strain: AB021422.
Synonym: "Bacterium woodsii" Smith 1911.
Etymology: N.L. gen. masc. n. woodsii, of Woods, named after A.F. Woods, an American plant pathologist.
Approved Lists reference: SKERMAN (V.B.D.), McGOWAN (V.) and SNEATH (P.H.A.) (editors): Approved Lists of Bacterial Names. Int. J. Syst. Bacteriol., 1980, 30, 225-420 (Approved Lists of Bacterial Names in IJSEM Online - Approved Lists of Bacterial Names Amended edition).
Original publication: 1 SMITH (E.F.): Bacteria in relation to plant diseases. Carnegie Institute, Washington, 1911, 2, 1-368. 2 STEVENS (F.L.) (ed.): Plant Disease Fungi, MacMillan Co., New York, 1925.

¤ Burkholderia andropogonis (Smith 1911) Gillis et al. 1995.
Reference: COENYE (T.), LAEVENS (S.), GILLIS (M.) and VANDAMME (P.): Genotypic and chemotaxonomic evidence for the reclassification of Pseudomonas woodsii (Smith 1911) Stevens 1925 as Burkholderia andropogonis (Smith 1911) Gillis et al. 1995. Int. J. Syst. Evol. Microbiol., 2001, 51, 183-185.
Original article in IJSEM Online

Note: Pseudomonas woodsii (Smith 1911) Stevens 1925 (Approved Lists 1980) and ¤ Burkholderia andropogonis (Smith 1911) Gillis et al. 1995 represent the same species. Based on Rule 42, Coenye et al. 2001 propose to retain the name ¤ Burkholderia andropogonis for this taxon.

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Pseudomonas xanthomarina Romanenko et al. 2005, sp. nov.
Type strain: (see also StrainInfo.net) CCUG 46543 = JCM 12468 = KMM 1447 = NRIC 0617.
Sequence accession no. (16S rRNA gene) for the type strain: AB176954.
Etymology: Gr. adj. xanthos, yellow; L. fem. adj. marina, of or belonging to the sea, marine; N.L. fem. adj. xanthomarina, yellow-pigmented bacteria from the sea.
Valid publication: Validation List no. 106. Int. J. Syst. Evol. Microbiol., 2005, 55, 2235-2238.
Effective publication: ROMANENKO (L.A.), UCHINO (M.), FALSEN (E.), LYSENKO (A.M.), ZHUKOVA (N.V.) and MIKHAILOV (V.V.): Pseudomonas xanthomarina sp. nov., a novel bacterium isolated from marine ascidian. J. Gen. Appl. Microbiol., 2005, 51, 65-71.
Validation List no. 106 in IJSEM Online - Effective publication Online

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Pseudomonas xiamenensis Lai and Shao 2008, sp. nov.
Type strain: (see also StrainInfo.net) C10-2 = CGMCC 1.6446 = JCM 13530 = MCCC 1A00089.
Sequence accession no. (16S rRNA gene) for the type strain: DQ088664.
Etymology: N.L. fem. adj. xiamenensis, of Xiamen, a district in Fujian, China, where the type strain was isolated.
Valid publication: LAI (Q.) and SHAO (Z.): Pseudomonas xiamenensis sp. nov., a denitrifying bacterium isolated from activated sludge. Int. J. Syst. Evol. Microbiol., 2008, 58, 1911-1915.
Original article in IJSEM Online

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Pseudomonas xinjiangensis Liu et al. 2009, sp. nov.
Type strain: (see also StrainInfo.net) S3-3 = CCTCC AB 207151 = NRRL B-51270.
Sequence accession no. (16S rRNA gene) for the type strain: EU286805.
Etymology: N.L. fem. adj. xinjiangensis, pertaining to Xinjiang, in north-west China, where the type strain was isolated.
Valid publication: LIU (M.), LUO (X.), ZHANG (L.), DAI (J.), WANG (Y.), TANG (Y.), LI (J.), SUN (T.) and FANG (C.): Pseudomonas xinjiangensis sp. nov., a moderately thermotolerant bacterium isolated from desert sand. Int. J. Syst. Evol. Microbiol., 2009, 59, 1286-1289.
Original article in IJSEM Online

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Pseudomonas yamanorum Arnau et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) 8H1=DSM 26522=CCUG 63249=LMG 27247.
Sequence accession no. (16S rRNA) for the type strain: EU557337.
Etymology: N.L. gen. pl. n. yamanorum, of the Yámanas, a tribe indigenous to the Ushuaia region, from where the type strain was isolated.
Source: Environmental - soil.
Valid publication: ARNAU, V. C. G., SÁNCHEZ, L. A. and DELGADO, O. D. 2015. Pseudomonas yamanorum sp. nov., a psychrotolerant bacterium isolated from a subantarctic environment. Int. J. Syst. Evol. Microbiol., 65, 424-431
Original article in IJSEM Online

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Pseudomonas zeshuii Feng et al. 2012, sp. nov.
Type strain: (see also StrainInfo.net) BY-1 = BY = ACCC 05688 = KACC 15471.
Sequence accession no. (16S rRNA gene) for the type strain: JN411093.
Etymology: N.L. masc. gen. n. zeshuii, of Ze-Shu, in honour of Ze-Shu Qian, a respected microbiologist, for his enormous contributions to promoting the development of soil microbiology in China.
Valid publication: FENG (Z.), ZHANG (J.), HUANG (X.), ZHANG (J.), CHEN (M.) and LI (S.): Pseudomonas zeshuii sp. nov., isolated from herbicide-contaminated soil. Int. J. Syst. Evol. Microbiol., 2012, 62, 2608-2612.
Original article in IJSEM Online

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Pseudomonas zhaodongensis Zhang et al. 2015, sp. nov.
Type strain: (see also StrainInfo.net) NEAU-ST5-21=ACCC 06362=DSM 27559.
Sequence accession no. (16S rRNA) for the type strain: JQ762275.
Etymology: N.L. fem. adj. zhaodongensis, pertaining to Zhaodong City, China, where the type strain was isolated.
Source: Environmental - soil.
Valid publication: ZHANG, L., PAN, Y., WANG, K., ZHANG, X., ZHANG, C., ZHANG, S., FU, X. and JIANG, J. 2015. Pseudomonas zhaodongensis sp. nov., isolated from saline and alkaline soils. Int. J. Syst. Evol. Microbiol., 65, 1022-1030.
Original article in IJSEM Online

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